Bifurcations for a predator-prey system with two delays

In this paper, a predator-prey system with two delays is investigated. By choosing the sum τ of two delays as a bifurcation parameter, we show that Hopf bifurcations can occur as τ crosses some critical values. By deriving the equation describing the flow on the center manifold, we can determine the direction of the Hopf bifurcations and the stability of the bifurcating periodic solutions. In addition, special attention is paid to the global continuation of local Hopf bifurcations. Using a global Hopf bifurcation result of [J. Wu, Symmetric functional differential equations and neural networks with memory, Trans. Amer. Math. Soc. 350 (1998) 4799-4838], we may show the global existence of periodic solutions.     

Hopf bifurcations to quasi-periodic solutions for the two-dimensional plane Poiseuille flow

This paper studies various Hopf bifurcations in the two-dimensional plane Poiseuille problem. For several values of the wavenumber α, we obtain the branch of periodic flows which are born at the Hopf bifurcation of the laminar flow. It is known that, taking α ≈ 1, the branch of periodic solutions has several Hopf bifurcations to quasi-periodic orbits. For the first bifurcation, calculations from other authors seem to indicate that the bifurcating quasi-periodic flows are stable and subcritical with respect to the Reynolds number, Re. By improving the precision of previous works we find that the bifurcating flows are unstable and supercritical with respect to Re. We have also analysed the second Hopf bifurcation of periodic orbits for several α, to find again quasi-periodic solutions with increasing Re. In this case the bifurcated solutions are stable to superharmonic disturbances for Re up to another new Hopf bifurcation to a family of stable 3-tori. The proposed numerical scheme is based on a full numerical integration of the Navier-Stokes equations, together with a division by 3 of their total dimension, and the use of a pseudo-Newton method on suitable Poincaré sections. The most intensive part of the computations has been performed in parallel. We believe that this methodology can also be applied to similar problems.     

Bifurcation analysis for a three-species predator-prey system with two delays

In this paper, a three-species predator-prey system with two delays is investigated. By choosing the sum τ of two delays as a bifurcation parameter, we first show that Hopf bifurcation at the positive equilibrium of the system can occur as τ crosses some critical values. Second, we obtain the formulae determining the direction of the Hopf bifurcations and the stability of the bifurcating periodic solutions by using the normal form theory and center manifold theorem. Finally, numerical simulations supporting our theoretical results are also included.     

Bifurcation analysis for a regulated logistic growth model

In this paper, we consider a regulated logistic growth model. We first consider the linear stability and the existence of a Hopf bifurcation. We show that Hopf bifurcations occur as the delay τ passes through critical values. Then, using the normal form theory and center manifold reduction, we derive the explicit algorithm determining the direction of Hopf bifurcations and the stability of the bifurcating periodic solutions. Finally, numerical simulation results are given to support the theoretical predictions.     

Dynamics of a competitive Lotka-Volterra system with three delays

In this paper, a competitive Lotka-Volterra system with three delays is investigated. By choosing the sum τ of three delays as a bifurcation parameter, we show that in the above system, Hopf bifurcation at the positive equilibrium can occur as τ crosses some critical values. And we obtain the formulae determining direction of Hopf bifurcation and stability of the bifurcating periodic solutions by using the normal form theory and center manifold theorem. Finally, numerical simulations supporting our theoretical results are also included.     

Hopf bifurcation in symmetric configuration of predator-prey-mutualist systems

In this paper we apply the equivariant degree method to study Hopf bifurcations in a system of differential equations describing a symmetric predator-prey-mutualist model with diffusive migration between interacting communities. A topological classification (according to symmetry types), of symmetric Hopf bifurcation in configurations of populations with D₈, D₁₂, A₄ and S₄ symmetries, is presented with estimation on minimal number of bifurcating branches of periodic solutions.     

Stability and Hopf bifurcation analysis of an eco-epidemic model with a stage structure

In this paper, an eco-epidemiological model with a stage structure is considered. The asymptotical stability of the five equilibria, the existence of stability switches about positive equilibrium, is investigated. It is found that Hopf bifurcation occurs when the delay τ passes though a critical value. Some explicit formulae determining the stability and the direction of the Hopf bifurcation periodic solutions bifurcating from Hopf bifurcations are obtained by using the normal form theory and center manifold theory. Some numerical simulations for justifying the theoretical analysis are also provided. Finally, biological explanations and main conclusions are given.     

The effect of time delay on the dynamics of an SEIR model with nonlinear incidence

In this paper, a SEIR epidemic model with nonlinear incidence rate and time delay is investigated in three cases. The local stability of an endemic equilibrium and a disease-free equilibrium are discussed using stability theory of delay differential equations. The conditions that guarantee the asymptotic stability of corresponding steady-states are investigated. The results show that the introduction of a time delay in the transmission term can destabilize the system and periodic solutions can arise through Hopf bifurcation when using the time delay as a bifurcation parameter. Applying the normal form theory and center manifold argument, the explicit formulas determining the properties of the bifurcating periodic solution are derived. In addition, the effect of the inhibitory effect on the properties of the bifurcating periodic solutions is studied. Numerical simulations are provided in order to illustrate the theoretical results and to gain further insight into the behaviors of delayed systems.     

Hopf bifurcation in a three-species system with delays

A kind of three-species system with Holling II functional response and two delays is introduced. Its local stability and the existence of Hopf bifurcation are demonstrated by analyzing the associated characteristic equation. By using the normal form method and center manifold theorem, explicit formulas to determine the direction of the Hopf bifurcation and the stability of bifurcating periodic solution are also obtained. In addition, the global existence results of periodic solutions bifurcating from Hopf bifurcations are established by using a global Hopf bifurcation result. Numerical simulation results are also given to support our theoretical predictions.     

Local Hopf bifurcation and global periodic solutions in a delayed predator-prey system

We consider a delayed predator-prey system. We first consider the existence of local Hopf bifurcations, and then derive explicit formulas which enable us to determine the stability and the direction of periodic solutions bifurcating from Hopf bifurcations, using the normal form theory and center manifold argument. Special attention is paid to the global existence of periodic solutions bifurcating from Hopf bifurcations. By using a global Hopf bifurcation result due to Wu [Trans. Amer. Math. Soc. 350 (1998) 4799], we show that the local Hopf bifurcation implies the global Hopf bifurcation after the second critical value of delay. Finally, several numerical simulations supporting the theoretical analysis are also given.     

Dynamics of a Delayed Predator-prey System with Stage Structure for Predator and Prey

In this paper, a predator-prey system with two discrete delays and stage structure for both the predator and the prey is investigated. The dynamical behaviors such as local stability and local Hopf bifurcation are analyzed by regarding the possible combinations of the two delays as bifurcating parameter. Some explicit formulae determining the direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions are derived by using the normal form method and the center manifold theory. Finally, numerical simulations are presented to support the theoretical analysis.     

Non-linear waves in a ring of neurons

^** Corresponding author. Email: shangjguo{at}etang.com In this paper, we study the effect of synaptic delay of signaltransmission on the pattern formation and some properties ofnon-linear waves in a ring of identical neurons. First, linearstability of the model is investigated by analyzing the associatedcharacteristic transcendental equation. Regarding the delayas a bifurcation parameter, we obtained the spontaneous bifurcationof multiple branches of periodic solutions and their spatio-temporalpatterns. Second, global continuation conditions for Hopf bifurcatingperiodic orbits are derived by using the equivariant degreetheory developed by Geba et al. and independently by Ize &Vignoli. Third, we show that the coincidence of these periodicsolutions is completely determined either by a scalar delaydifferential equation if the number of neurons is odd, or bya system of two coupled delay differential equations if thenumber of neurons is even. Fourth, we summarize some importantresults about the properties of Hopf bifurcating periodic orbits,including the direction of Hopf bifurcation, stability of theHopf bifurcating periodic orbits, and so on. Fifth, in an excitatoryring network, solutions of most initial conditions tend to stableequilibria, the boundary separating the basin of attractionof these stable equilibria contains all of periodic orbits andhomoclinic orbits. Finally, we discuss a trineuron network toillustrate the theoretical results obtained in this paper andconclude that these theoretical results are important to complementthe experimental and numerical observations made in living neuronssystems and artificial neural networks, in order to understandthe mechanisms underlying the system dynamics better.     

含两时滞捕食-被捕食系统的稳定性及分歧

本文研究一类含两相异时滞的捕食－被捕食系统的稳定性及分歧。首先，我们讨论两相异时滞对系统唯一正平衡点的稳定性的影响，通过对系数与时滞有关的特征方程的分析，建立了一种稳定性判别性。其次，将一个时滞看成分歧参数，而另一个看作固定参数，我们证明了该系统具有HOPF分歧特性。最后，我们讨论了分歧解的稳定性。     

Hopf bifurcation for neutral functional differential equations

In this paper, we extend the computation of the properties of Hopf bifurcation, such as the direction of bifurcation and stability of bifurcating periodic solutions, of DDE introduced by Kazarinoff et al. [N.D. Kazarinoff, P. van den Driessche, Y.H. Wan, Hopf bifurcation and stability of periodic solutions of differential–difference and integro-differential equations, J. Inst. Math. Appl. 21 (1978) 461–477] to a kind of neutral functional differential equation (NFDE). As an example, a neutral delay logistic differential equation is considered, and the explicit formulas for determining the direction of bifurcation and the stability of bifurcating periodic solutions are derived. Finally, some numerical simulations are carried out to support the analytic results.     

Stability of Periodic Solutions Generated by Hopf Points Emanating from a Z＿2－symmetry－breaking Takens－Bogdanov Point

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Stability and Hopf bifurcation for a regulated logistic growth model with discrete and distributed delays

In this paper, we investigate the stability and Hopf bifurcation of a new regulated logistic growth with discrete and distributed delays. By choosing the discrete delay τ as a bifurcation parameter, we prove that the system is locally asymptotically stable in a range of the delay and Hopf bifurcation occurs as τ crosses a critical value. Furthermore, explicit algorithm for determining the direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions is derived by normal form theorem and center manifold argument. Finally, an illustrative example is also given to support the theoretical results.     

Hopf bifurcation and multiple periodic solutions in Lotka–Volterra systems with symmetries

The purpose of this paper is to study Hopf bifurcations in a delayed Lotka–Volterra system with dihedral symmetry. By treating the response delay as bifurcation parameter and employing equivariant degree method, we obtain the existence of multiple branches of nonconstant periodic solutions through a local Hopf bifurcation around an equilibrium. We find that competing coefficients and the response delay in the system can affect the spatio-temporal patterns of bifurcating periodic solutions. According to their symmetric properties, a topological classification is given for these periodic solutions. Furthermore, an estimation is presented on minimal number of bifurcating branches. These theoretical results are helpful to better understand the complex dynamics induced by response delays and symmetries in Lotka–Volterra systems.     

Stability and Hopf bifurcation in a delayed competition system

In this paper, Hopf bifurcation for two-species Lotka–Volterra competition systems with delay dependence is investigated. By choosing the delay as a bifurcation parameter, we prove that the system is stable over a range of the delay and beyond that it is unstable in the limit cycle form, i.e., there are periodic solutions bifurcating out from the positive equilibrium. Our results show that a stable competition system can be destabilized by the introduction of a maturation delay parameter. Further, an explicit algorithm for determining the direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions is derived by using the theory of normal forms and center manifolds, and numerical simulations supporting the theoretical analysis are also given.     

Two new approaches to computing Hopf bifurcation problems

We consider the computation of Hopf bifurcation for ordinary differential equations. Two new extended systems are given for the calculation of Hopf bifurcation problems: the first is composed of differential-algebraic equations with index 1, the other consists of differential equations by using a symmetry inherited from the autonomous system of ordinary differential equations. Both methods are especially suitable for calculating bifurcating periodic solutions since they transform the Hopf bifurcation problem into regular nonlinear boundary value problems which are very easy to implement. The bifurcation solutions become isolated solutions of the extended system so that our methods work both in the subcritical and supercritical case. The extended systems are based on an additional parameter ε; practical experience shows that one gets convergence for ε sufficiently large so that a substantial part of the bifurcating branch can be computed. The two methods are illustrated by numerical examples and compared with other procedures.     

具有捕食者相互残杀项时滞系统的Hopf分支

研究了具有捕食者相互残杀项的时滞系统的Hopf分支,通过选择时滞作为一个分支参数,研究了正平衡点的稳定性和正周期解的Hopf分支.而且通过应用规范型和中心流形的理论,得出了确定分支方向的明确的算法.