Stability analysis in a diffusional immunosuppressive infection model with delayed antiviral immune response

In this paper, the diffusion is introduced to an immunosuppressive infection model with delayed antiviral immune response. The direction and stability of Hopf bifurcation are effected by time delay, in the absence of which the positive equilibrium is locally asymptotically stable by means of analyzing eigenvalue spectrum; however, when the time delay increases beyond a threshold, the positive equilibrium loses its stability via the Hopf bifurcation. The stability and direction of the Hopf bifurcation is investigated with the norm form and the center manifold theory. The stability of the Hopf bifurcation leads to the emergence of spatial spiral patterns. Numerical calculations are performed to illustrate our theoretical results. Copyright © 2017 John Wiley & Sons, Ltd.     

Modeling on an ecological food chain with recycling

We propose two nutrient-phytoplankton models with instantaneous and time delayed recycling, investigate the dynamics and examine the responses to model complexities. Instead of the familiar specific uptake rate and growth rate functions, we assume only that the nutrient uptake and phytoplankton growth rate functions are positive, increasing and bounded above. We use geometrical and analytical methods to find conditions for the existence of none, one, or at most two positive steady states and analyze the stability properties of each of these equilibria. With the variation of parameters, the system may lose its stability and bifurcation may occur. We study the occurrence of Hopf bifurcation and the possibility of stability switching. Numerical simulations illustrate the analytical results and provide further insight into the dynamics of the models, biological interpretations are given.     

带有非线性收获和妊娠时滞的微分代数捕食者-食饵系统的稳定性及Hopf分支

本文研究了一类同时带有非线性食饵收获和捕食者妊娠时滞的微分代数捕食者-食饵系统的稳定性及Hopf分支问题.利用了分支理论和稳定性理论,以捕食者妊娠时滞作为系统的分支参数,获得了所提出的新系统在正平衡点处系统稳定性的相关判据条件和Hopf分支的产生条件.推广了一般带有线性收获和时滞的微分代数捕食者-食饵系统的结论.     

Stability Analysis of an Enterprise Competitive Model with Time Delay

A three-dimensional enterprise competitive model with time delay is considered. Where the delay is regarded as bifurcation parameters. By analyzing the corresponding characteristic equation of positive equilibrium,the local stability of positive equilibrium is regarded. By using the normal form method and center manifold theorem, we give the formula for determining the direction of the Hopf bifurcation and the stability of bifurcating periodic solutions. Numerical simulations are shown to illustrate the obtained results.     

一类带无选择性捕获和时滞的捕食食饵系统的Hopf分支分析

本文主要研究了一个改进的带时滞和无选择捕获函数的捕食-食饵生态经济系统的稳定性和Hopf分支.利用微分代数系统的稳定性理论和分支理论,得到了系统正平衡点稳定性的条件,以及当时滞τ作为分支参数时系统产生Hopf分支的条件.对Leslie-Gower捕食-食饵模型进行了一定程度的完善,使得建立的模型更符合实际情况,因此得到的结论也更加科学.     

Stability and Hopf bifurcation analysis on a spruce-budworm model with delay

In this paper, the dynamics of a spruce-budworm model with delay is investigated. We show that there exists Hopf bifurcation at the positive equilibrium as the delay increases. Some sufficient conditions for the existence of Hopf bifurcation are obtained by investigating the associated characteristic equation. By using the theory of normal form and center manifold, explicit expression for determining the direction of Hopf bifurcations and the stability of bifurcating periodic solutions are presented.     

一类具有时滞的捕食——食饵系统的稳定性与Hopf分支

研究了一类具有时滞的捕食—食饵系统,通过分析正平衡点处的特征方程,讨论了系统正平衡点的稳定性;以时滞作为分支参数,应用Hopf分支理论,得到了系统存在Hopf分支的充分条件.     

Stability and bifurcations for the chronic state in Marchuk's model of an immune system

In the paper we present known and new results concerning stability and the Hopf bifurcation for the positive steady state describing a chronic disease in Marchuk's model of an immune system. We describe conditions guaranteeing local stability or instability of this state in a general case and for very strong immune system. We compare these results with the results known in the literature. We show that the positive steady state can be stable only for very specific parameter values. Stability analysis is illustrated by Mikhailov's hodographs and numerical simulations. Conditions for the Hopf bifurcation and stability of arising periodic orbit are also studied. These conditions are checked for arbitrary chosen realistic parameter values. Numerical examples of arising due to the Hopf bifurcation periodic solutions are presented.     

Bifurcation behaviours in a delayed three-species food-chain model with Holling type-II functional response

This article is concerned with the local stability of a positive equilibrium and the Hopf bifurcation of a delayed three-species food-chain system with the Holling type-II functional response. Some new sufficient conditions ensuring the local stability of a positive equilibrium and the existence of Hopf bifurcation for the system are established. Some explicit formulae are derived to determine the direction of bifurcations and the stability of bifurcating periodic solutions using the normal form theory and the centre manifold theory. Numerical simulations for justifying the theoretical analysis are also provided. Finally, biological explanations and main conclusions are included.     

Spatial resonance and Turing–Hopf bifurcations in the Gierer–Meinhardt model

Gierer–Meinhardt system as a molecularly plausible model has been proposed to formalize the observation for pattern formation. In this paper, the Gierer–Meinhardt model without the saturating term is considered. By the linear stability analysis, we not only give out the conditions ensuring the stability and Turing instability of the positive equilibrium but also find the parameter values where possible Turing–Hopf and spatial resonance bifurcation can occur. Then we develop the general algorithm for the calculations of normal form associated with codimension-2 spatial resonance bifurcation to better understand the dynamics neighboring of the bifurcating point. The spatial resonance bifurcation reveals the interaction of two steady state solutions with different modes. Numerical simulations are employed to illustrate the theoretical results for both the Turing–Hopf bifurcation and spatial resonance bifurcation. Some expected solutions including stable spatially inhomogeneous periodic solutions and coexisting stable spatially steady state solutions evolve from Turing–Hopf bifurcation and spatial resonance bifurcation respectively.     

Dynamics of a delayed predator–prey model with predator migration

Based on the availability of prey and a simple predator–prey model, we propose a delayed predator–prey model with predator migration to describe biological control. We first study the existence and stability of equilibria. It turns out that backward bifurcation occurs with the migration rate as bifurcation parameter. The stability of the trivial equilibrium and the boundary equilibrium is delay-independent. However, the stability of the positive equilibrium may be delay-dependent. Moreover, delay can switch the stability of the positive equilibrium. When the positive equilibrium loses stability, Hopf bifurcation can occur. The direction and stability of Hopf bifurcation is derived by applying the center manifold method and the normal form theory. The main theoretical results are illustrated with numerical simulations.     

Dynamics of a ratio‐dependent stage‐structured predator‐prey model with delay

In this paper, we investigate the dynamics of a time‐delay ratio‐dependent predator‐prey model with stage structure for the predator. This predator‐prey system conforms to the realistically biological environment. The existence and stability of the positive equilibrium are thoroughly analyzed, and the sufficient and necessary conditions for the stability and instability of the positive equilibrium are obtained for the case without delay. Then, the influence of delay on the dynamics of the system is investigated using the geometric criterion developed by Beretta and Kuang. 26 We show that the positive steady state can be destabilized through a Hopf bifurcation and there exist stability switches under some conditions. The formulas determining the direction and the stability of Hopf bifurcations are explicitly derived by using the center manifold reduction and normal form theory. Finally, some numerical simulations are performed to illustrate and expand our theoretical results.     

Stability and Hopf bifurcation in a ratio-dependent predator–prey system with stage structure

A ratio-dependent predator–prey model with stage structure for the predator and time delay due to the gestation of the predator is investigated. By analyzing the characteristic equations, the local stability of a positive equilibrium and a boundary equilibrium is discussed, respectively. Further, it is proved that the system undergoes a Hopf bifurcation at the positive equilibrium when τ = τ₀. By using an iteration technique, sufficient conditions are derived for the global attractivity of the positive equilibrium. By comparison arguments, sufficient conditions are obtained for the global stability of the boundary equilibrium. Numerical simulations are carried out to illustrate the main results.     

Hopf bifurcations in a predator–prey system with a discrete delay and a distributed delay

A delayed Lotka–Volterra two-species predator–prey system with discrete hunting delay and distributed maturation delay for the predator population described by an integral with a strong delay kernel is considered. By linearizing the system at the positive equilibrium and analyzing the associated characteristic equation, the asymptotic stability of the positive equilibrium is investigated and Hopf bifurcations are demonstrated. It is found that under suitable conditions on the parameters the positive equilibrium is asymptotically stable when the hunting delay is less than a certain critical value and unstable when the hunting delay is greater than this critical value. Meanwhile, according to the Hopf bifurcation theorem for functional differential equations (FDEs), we find that the system can also undergo a Hopf bifurcation of nonconstant periodic solution at the positive equilibrium when the hunting delay crosses through a sequence of critical values. In particular, by applying the normal form theory and the center manifold reduction for FDEs, an explicit algorithm determining the direction of Hopf bifurcations and the stability of bifurcating periodic solutions occurring through Hopf bifurcations is given. Finally, to verify our theoretical predictions, some numerical simulations are also included at the end of this paper.     

Dynamical behavior of a delay virus dynamics model with CTL immune response

In this paper, the dynamical behavior of a virus dynamics model with CTL immune response and time delay is studied. Time delay is used to describe the time between the infected cell and the emission of viral particles on a cellular level. The effect of time delay on stability of the equilibria of the CTL immune response model has been studied and sufficient criteria for local asymptotic stability of the disease-free equilibrium, immune-free equilibrium and endemic equilibrium and global asymptotic stability of the disease-free equilibrium are given. Some conditions for Hopf bifurcation around immune-free equilibrium and endemic equilibrium to occur are also obtained by using the time delay as a bifurcation parameter. Numerical simulation with some hypothetical sets of data has been done to support the analytical findings.     

Bifurcation analysis of a modified Holling-Tanner predator-prey model with time delay

In this paper, a modified Holling-Tanner predator-prey model with time delay is considered. By regarding the delay as the bifurcation parameter, the local asymptotic stability of the positive equilibrium is investigated. Meanwhile, we find that the system can also undergo a Hopf bifurcation of nonconstant periodic solution at the positive equilibrium when the delay crosses through a sequence of critical values. In particular, we study the direction of Hopf bifurcation and the stability of bifurcated periodic solutions, an explicit algorithm is given by applying the normal form theory and the center manifold reduction for functional differential equations. Finally, numerical simulations supporting the theoretical analysis are also included.     

The delay effect on reaction–diffusion equations

In this work we study a reaction–diffusion problem with delay and we make an analysis of the stability of solutions by means of bifurcation theory. We take the delay constant as a parameter. Special conditions on the vector field assure existence of a spatially nonconstant positive equilibrium U_k , which is stable for small values of the delay. An increase of the delay destabilizes the equilibrium of U_k and leads to super or subcritical Hopf bifurcation.     

一类具有时滞的云杉蚜虫种群模型的Hopf分岔分析

研究了一类具有时滞的云杉蚜虫种群阶段结构模型的动力学行为.首先,讨论了模型正平衡点的存在唯一性,并分析了该平衡点的局部稳定性和出现Hopf分岔的充分条件;其次,利用中心流形定理和正规形理论,讨论了分岔周期解的稳定性及方向;最后,通过数值模拟验证了相关结论的正确性.该文所得结论具有广泛的实际应用价值.     

Stability and Hopf bifurcation analysis in a three-level food chain system with delay

A class of three level food chain system is studied. With the theory of delay equations and Hopf bifurcation, the conditions of the positive equilibrium undergoing Hopf bifurcation is given regarding τ as the parameter. The stability and direction of Hopf bifurcation are determined by applying the normal form theory and the center manifold argument, and numerical simulations are performed to illustrate the analytical results.     

Stability and bifurcation in a harmonic oscillator with delays

We consider a harmonic oscillator with delays. Linear stability is investigated by analyzing the associated characteristic transcendental equation. The bifurcation analysis of the equation shows that Hopf bifurcation can occur as the delay τ (taken as a parameter) crosses some critical values. The direction and stability of the Hopf bifurcation are considered by using the normal form theory due to Faria and Magalhães. An example is given to explain the results. Numerical simulations support our results.