Effect of prey refuge on a harvested predator–prey model with generalized functional response

A predator–prey model with generalized response function incorporating a prey refuge and independent harvesting in each species are studied by using the analytical approach. A constant proportion of prey using refuges is considered. We will evaluate the effects with regard to the local stability of equilibria, the equilibrium density values and the long-term dynamics of the interacting populations. Some numerical simulations are carried out.     

具有阶段结构且食饵有避难所的模型分析

研究了捕食者具有阶段结构且食饵有避难所的非自治捕食系统.利用Lyapunov函数方法得到了系统持续生存的条件,以及在一定条件下存在唯一全局渐进稳定的周期正解.对于更广泛的概周期现象,也得到了存在唯一全局渐进稳定的概周期正解的充分条件.     

Consequences of refuge and diffusion in a spatiotemporal predator–prey model

In this investigation, we offer and examine a predator–prey interacting model with prey refuge in proportion to both the species and Beddington–DeAngelis functional response. We first prove the well-posedness of the temporal and spatiotemporal models which are restricted in a positive invariant region. Then for the temporal model, we analyse its temporal dynamics including uniform boundedness, permanence, stability of all feasible non-negative equilibria and show that refugia can induce periodic oscillation via Hopf bifurcation around the unique positive equilibrium; for the spatiotemporal model, we not only investigate its permanence, stability of non-negative constant steady states and Turing instability but also study the existence and non-existence of non-constant positive steady states by Leray–Schauder degree theory. The key observation is that the coefficient of refuge cooperates a significant part in modifying the dynamics of the current system and mediates the population permanence, stability of coexisting equilibrium and even the Turing instability parameter space. Finally, general numerical simulation consequences are given to illustrate the validity of the theoretical results. Through numerical simulations, one observes that the model dynamics shows prey refugia and self-diffusion control spatiotemporal pattern growth to spots, stripe–spot mixtures and stripes reproduction. The outcomes assign that the dynamics of the model with prey refuge is not simple, but rich and complex. Additionally, numerical simulations show that the other model parameters have an important effect on species’ spatially inhomogeneous distribution, which results in the formation of spots pattern, mixture of spots and stripes pattern, mixture of spots, stripes and rings pattern and anti-spot pattern. This may improve the model dynamics of the prey refuge on the reaction–diffusion predator–prey system.     

具有食饵避难的Leslie-Gower捕食系统最优收获分析

研究了一类具有食饵避难的Leslie-Gower捕食与被捕食系统收获模型,利用Hurwitz判据,得到了正平衡点局部渐近稳定,进一步构造了适当的Lyapunov函数,证明了正平衡点的全局渐近稳定性.并且在捕获努力量假说下,对发生食饵避难的两种群同时捕获,考虑了生态经济平衡点的存在性和利用Pontryagin最大值原理对两种群进行最优收获,得到当贴现率为零时,既保持了生态平衡,又使得在渔业开发过程中取得最大经济利益.     

一类具第三类功能反应且食饵具有避难所的捕食系统的分析

研究了一类具第三类功能反应且食饵具有避难所的非自治捕食系统.利用Lyapunov函数方法得到了系统持续生存的条件,以及在一定条件下,系统存在全局渐进稳定的周期正解.对于更广泛的概周期现象,也得到了存在唯一全局渐进稳定的概周期正解的充分条件.     

Hopf bifurcation in a diffusive predator-prey model with herd behavior and prey harvesting

In this paper, the dynamics of a diffusive delayed predator-prey model with herd behavior and prey harvesting subject to the homogeneous Neumann boundary condition is considered. Firstly, choosing the harvesting term as a bifurcation parameter, then we obtain the existence and the stability of the equilibrium by analyzing the distribution of the roots of associated characteristic equation. Secondly, time delay is regarding as a bifurcation parameter, and the use of the normal form theory and center manifold theorem, the existence, stability and direction of bifurcating periodic solutions are all demonstrated detailly. Finally, summarizing some numerical simulations to illustrate the theoretical analysis.     

On Nonautonomous Prey predator Patchy System

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A ratio‐dependent predator–prey model with stage structure and optimal harvesting policy

In this paper, a ratio‐dependent predator–prey model with stage structure and harvesting is investigated. Mathematical analyses of the model equations with regard to boundedness of solutions, nature of equilibria, permanence and stability are performed. By constructing appropriate Lyapunov functions, a set of easily verifiable sufficient conditions are obtained for the global asymptotic stability of nonnegative equilibria of the model. The existence possibilities of bioeconomic equilibria have been examined. An optimal harvesting policy is also given by using Pontryagin's maximal principle. Copyright © 2007 John Wiley & Sons, Ltd.     

Chaos and Hopf bifurcation analysis for a two species predator–prey system with prey refuge and diffusion

In this paper, a delayed predator–prey model with Holling type II functional response incorporating a constant prey refuge and diffusion is considered. By analyzing the characteristic equation of linearized system corresponding to the model, we study the local asymptotic stability of the positive equilibrium of the system. By choosing the time delay due to gestation as a bifurcation parameter, the existence of Hopf bifurcations at the positive equilibrium is established. By applying the normal form and the center manifold theory, an explicit algorithm to determine the direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions is derived. Further, an example is presented to illustrate our main results. Finally, recurring to the numerical method, the influences of impulsive perturbations on the dynamics of the system are also investigated.     

Dynamics of a ratio-dependent eco-epidemiological system with prey harvesting

In this article, we study a ratio-dependent eco-epidemiological system where prey population is subjected to harvesting. Mathematical results like positive invariance, boundedness, stability of equilibria, and permanence of the system have been established. The dynamics of zero equilibria have been thoroughly investigated to find out conditions on the system parameters such that trajectories starting from the domain of interest can reach a zero equilibrium following any fixed direction. We have also studied suitable conditions for non-existence of a periodic solution around the interior equilibrium. Computer simulations have been carried out to illustrate different analytical findings.     

ALMOST PERIODIC SOLUTION OF A NONAUTONOMOUS MODIFIED LESLIE-GOWER PREDATOR-PREY MODEL WITH NONMONOTONIC FUNCTIONAL RESPONSE AND A PREY REFUGE

A nonautonomous modified Leslie-Gower predator-prey model with nonmonotonic functional response and a prey refuge is proposed and studied in this paper. Sufficient conditions which guarantee the permanence, extinction of the prey species and the global stability of the system are obtained, respectively. Also, by constructing a suitable Lyapunov function, some sufficient conditions are obtained for the existence of a unique globally attractive positive almost periodic solution of this model. Our results indicate that the prey refuge has positive effect on the coexistence of the species. Examples together with their numeric simulation show the feasibility of our main results.     

Hopf bifurcation analysis for a delayed predator-prey system with a prey refuge and selective harvesting

In this paper, a delayed predator-prey system with Holling type III functional response incorporating a prey refuge and selective harvesting is considered. By analyzing the corresponding characteristic equations, the conditions for the local stability and existence of Hopf bifurcation for the system are obtained, respectively. By utilizing normal form method and center manifold theorem, the explicit formulas which determine the direction of Hopf bifurcation and the stability of bifurcating period solutions are derived. Finally, numerical simulations supporting the theoretical analysis are given.     

Dynamics of transitions in population interactions

A two-species model with transitions between population interactions is studied. Rich dynamics is observed as the number and quality of equilibria change when model parameters and functional responses vary. Existence and stability of equilibria and nonexistence of periodic solutions are established, existence of some bifurcation phenomena are analytically and numerically studied, explicit threshold values are computed to determine the kind of interaction (mutualism, competition, host-parasite) between the species, and several numerical examples are provided to illustrate the main results in this work.     

Complex dynamics of a diffusive predator–prey model with strong Allee effect and threshold harvesting

In this paper, complex dynamics of a diffusive predator–prey model is investigated, where the prey is subject to strong Allee effect and threshold harvesting. The existence and stability of nonnegative constant steady state solutions are discussed. The existence and nonexistence of nonconstant positive steady state solutions are analyzed to identify the ranges of parameters of pattern formation. Spatially homogeneous and nonhomogeneous Hopf bifurcation and discontinuous Hopf bifurcation are proved. These results show that the introduction of strong Allee effect and threshold harvesting increases the system spatiotemporal complexity. Finally, numerical simulations are presented to validate the theoretical results.     

Spatiotemporal dynamics of a Leslie–Gower predator–prey model incorporating a prey refuge

In this paper, we investigate the spatiotemporal dynamics of a two-dimensional predator–prey model, which is based on a modified version of the Leslie–Gower scheme incorporating a prey refuge. We establish a Lyapunov function to prove the global stability of the equilibria with diffusion and determine the Turing space in the spatial domain. Furthermore, we perform a series of numerical simulations and find that the model dynamics exhibits complex Turing pattern replication: stripes, cold/hot spots-stripes coexistence and cold/hot spots patterns. The results indicate that the effect of the prey refuge for pattern formation is tremendous. This may enrich the dynamics of the effect of refuge on the predator-prey systems.     

On non-selective harvesting of a multispecies fishery

The present paper deals with the problem of non-selective harvesting of a prey-predator system in which both the prey and the predator species obey the law of logistic growth and each predators functional response to the prey approaches a constant as the prey population increases. Boundedness of the exploited system is examined. The existence of its steady states and their stability are studied using eigenvalue analysis. The existence of bionomic equilibria has been considered. The problem of determining the optimal harvest policy is then solved by using Pontryagin's maximal principle. Finally, some numerical examples are given to illustrate the results.     

Saddle-node-Hopf bifurcation in a modified Leslie–Gower predator-prey model with time-delay and prey harvesting

From the saddle-node-Hopf bifurcation point of view, this paper considers a modified Leslie–Gower predator-prey model with time delay and the Michaelis–Menten type prey harvesting. Firstly, we discuss the stability of the equilibria, obtain the critical conditions for the saddle-node-Hopf bifurcation, and give the completion bifurcation set by calculating the universal unfoldings near the saddle-node-Hopf bifurcation point by using the normal form theory and center manifold theorem. Then we derive the parameter conditions for the existence of monostable coexistence equilibrium and the parameter regions in which both the prey-extinction and the coexistence equilibrium (or coexistence periodic or quasi-periodic solutions) are simultaneously stabilized. We also investigate the heteroclinic bifurcation, and describe the phenomenon that the periodic behavior disappears as through the heteroclinic bifurcation. Finally, some numerical simulations are performed to support our analytic results.     

A Leslie–Gower predator–prey model with disease in prey incorporating a prey refuge

In this paper, a predator–prey Leslie–Gower model with disease in prey has been developed. The total population has been divided into three classes, namely susceptible prey, infected prey and predator population. We have also incorporated an infected prey refuge in the model. We have studied the positivity and boundedness of the solutions of the system and analyzed the existence of various equilibrium points and stability of the system at those equilibrium points. We have also discussed the influence of the infected prey refuge on each population density. It is observed that a Hopf bifurcation may occur about the interior equilibrium taking refuge parameter as bifurcation parameter. Our analytical findings are illustrated through computer simulation using MATLAB, which show the reliability of our model from the eco-epidemiological point of view.     

Dynamics analysis of a reaction–diffusion system with Beddington–DeAngelis functional response and strong Allee effect

In this paper, the dynamics of a diffusive predator–prey model with modified Leslie–Gower term and strong Allee effect on prey under homogeneous Neumann boundary condition is considered. Firstly, we obtain the qualitative properties of the system including the existence of the global positive solution and the local and global asymptotical stability of the constant equilibria. In addition, we investigate a priori estimate and the nonexistence of nonconstant positive steady state solutions. Finally, we establish the existence and local structure of steady state patterns and time-periodic patterns for the system.     

Dynamical behavior analysis of a two-dimensional discrete predator-prey model with prey refuge and fear factor

This paper investigates the dynamics of an improved discrete Leslie-Gower predator-prey model with prey refuge and fear factor. First, a discrete Leslie-Gower predator-prey model with prey refuge and fear factor has been introduced. Then, the existence and stability of fixed points of the model are analyzed. Next, the bifurcation behaviors are discussed, both flip bifurcation and Neimark-Sacker bifurcation have been studied. Finally, some simulations are given to show the effectiveness of the theoretical results.