A bifurcation theorem for nonlinear matrix models of population dynamics

ABSTRACT

We prove a general theorem for nonlinear matrix models of the type used in structured population dynamics that describes the bifurcation that occurs when the extinction equilibrium destabilizes as a model parameter is varied. The existence of a bifurcating continuum of positive equilibria is established, and their local stability is related to the direction of bifurcation. Our theorem generalizes existing theorems found in the literature in two ways. First, it allows for a general appearance of the bifurcation parameter (existing theorems require the parameter to appear linearly). This significantly widens the applicability of the theorem to population models. Second, our theorem describes circumstances in which a backward bifurcation can produce stable positive equilibria (existing theorems allow for stability only when the bifurcation is forward). The signs of two diagnostic quantities determine the stability of the bifurcating equilibrium and the direction of bifurcation. We give examples that illustrate these features.     

Stability and Hopf bifurcations in a business cycle model with delay

In this paper, a business cycle model with discrete delay is considered. We first investigate the stability of the equilibrium and the existence of Hopf bifurcations, and then the direction and the stability criteria of the bifurcating periodic solutions are obtained by the normal form theory and the center manifold theorem. This research has an important theoretical value as well as practical meaning.     

Hopf bifurcation in REM algorithm with communication delay

The purpose of this paper is to study bifurcation of an Internet congestion control algorithm, namely REM (Random Exponential Marking) algorithm, with communication delay. By choosing the delay constant as a bifurcation parameter, we prove that REM algorithm exhibits Hopf bifurcation. The formulas for determining the direction of the Hopf bifurcation and the stability of bifurcating periodic solutions are obtained by applying the center manifold theorem and the normal form theory. Finally, a numerical simulation is present to verify the theoretical results.     

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In this paper, we first study the strong law of large numbers for the frequencies of occurrence of random ordered couples of states for nonhomogeneous bifurcating Markov chains indexed by a binary tree. Then the strong law of large numbers are studied for functions of the nonhomogeneous bifurcating Markov chains indexed by a binary tree. As a corollary, we obtain the shannon-McMillan theorem for these Markov chains with finite state space.     

Stability and Hopf bifurcation analysis on a four-neuron BAM neural network with distributed delays

In this paper, a four-neuron BAM neural network with distributed delays is considered, where kernels are chosen as weak kernels. Its dynamics is studied in terms of local stability analysis and Hopf bifurcation analysis. By choosing the average delay as a bifurcation parameter and analyzing the associated characteristic equation, Hopf bifurcation occurs when the bifurcation parameter passes through some exceptive values. The stability of bifurcating periodic solutions and a formula for determining the direction of Hopf bifurcation are determined by applying the normal form theory and the center manifold theorem. Finally, numerical simulation results are given to validate the theorem obtained.     

Stability Switches, Hopf Bifurcations, and Spatio-temporal Patterns in a Delayed Neural Model with Bidirectional Coupling

The dynamical behavior of a delayed neural network with bi-directional coupling is investigated by taking the delay as the bifurcating parameter. Some parameter regions are given for conditional/absolute stability and Hopf bifurcations by using the theory of functional differential equations. As the propagation time delay in the coupling varies, stability switches for the trivial solution are found. Conditions ensuring the stability and direction of the Hopf bifurcation are determined by applying the normal form theory and the center manifold theorem. We also discuss the spatio-temporal patterns of bifurcating periodic oscillations by using the symmetric bifurcation theory of delay differential equations combined with representation theory of Lie groups. In particular, we obtain that the spatio-temporal patterns of bifurcating periodic oscillations will alternate according to the change of the propagation time delay in the coupling, i.e., different ranges of delays correspond to different patterns of neural activities. Numerical simulations are given to illustrate the obtained results and show the existence of bursts in some interval of the time for large enough delay.     

Stability and bifurcation of a Cohen-Grossberg neural network with discrete delays

A Cohen-Grossberg neural network with discrete delays is investigated in this paper. The qualitative analysis is given for the system and it is found that the system undergoes a sequence of Hopf bifurcations by choosing the discrete time delay as a bifurcation parameter. Moreover, by applying the normal form theory and the center manifold theorem, the direction of the Hopf bifurcation and the stability of bifurcating periodic solutions are obtained. Numerical simulations are given to illustrate the obtained results.     

Stability and Hopf Bifurcation of a Type of Protein Synthesis System with Time Delay and Negative Feedback

This paper studied the stability and Hopf bifurcation of a type of protein synthesis system with time delay and negative feedback. Firstly, it is proved theoretically that the time delay, nonlinearity in the protein production and the cooperativity in the negative feedback are key factors to generate circadian oscillation; Taking time delay as a parameter, we obtained the critical value of the time delay that Hopf bifurcation generates. Secondly, based on the center manifold and normal form theorem, we derived the formulas for determining the stability of bifurcating periodic solutions and the supercritical or subcritical Hopf bifurcation. Finally, the matlab program is used to simulate the results.     

Dynamics of a congestion control model in a wireless access network

In this paper, a congestion control algorithm with heterogeneous delays in a wireless access network is considered. We regard the communication time delay as a bifurcating parameter to study the dynamical behaviors, i.e., local asymptotical stability, Hopf bifurcation and resonant codimension-two bifurcation. By analyzing the associated characteristic equation, the Hopf bifurcation occurs when the delay passes through a sequence of critical value. Furthermore, the direction and stability of the bifurcating periodic solutions are derived by applying the normal form theory and the center manifold theorem. In the meantime, the resonant codimension-two bifurcation is also found in this model. Some numerical examples are finally performed to verify the theoretical results.     

Hopf bifurcation in a three-species system with delays

A kind of three-species system with Holling II functional response and two delays is introduced. Its local stability and the existence of Hopf bifurcation are demonstrated by analyzing the associated characteristic equation. By using the normal form method and center manifold theorem, explicit formulas to determine the direction of the Hopf bifurcation and the stability of bifurcating periodic solution are also obtained. In addition, the global existence results of periodic solutions bifurcating from Hopf bifurcations are established by using a global Hopf bifurcation result. Numerical simulation results are also given to support our theoretical predictions.     

In “On invariant surfaces and bifurcation of periodic solutions of ordinary differential equations. Chapter II: Bifurcation-mapping method” by Robert J. Sacker

This article consists of Chapter 2 of the author's 1964 Dissertation bearing the same title and published as Courant Institute report IMM-NYU 333, available in full on the author's personal web site. Chapter 2 consists of the first complete proof of what has come to be known as the Neimark-Sacker bifurcation theorem. It includes the reduction to normal form using weighted monomials which precludes using the Center Manifold theorem which was not known to the author and was published the same year as the dissertation and called “a Reduction Principle” by V. Pliss and later named Center Manifold theorem by A. Kelley (see previous article for citations). After reduction to normal form, the resulting functional equation for the bifurcating invariant curve is solved in detail. Stand-alone lemmas provide solutions of linear functional equations, a-priori estimates and interpolation inequalities between derivatives in the sup norm.     

Direction and stability of bifurcating periodic solutions of a chemostat model with two distributed delays

We have considered a chemostat model with two distributed delays in a recent paper [Chaos, Solitons & Fractals 2004;20:995–1004], where, using the average time delay corresponding to the growth response as a bifurcation parameter, it is proven that the model undergoes Hopf bifurcations for two weak kernels. This article is a sequel to the previous work. The direction and stability of the bifurcating periodic solutions are determined by the normal form theory and the center manifold theorem. The results are consistent with the numerical results in [Chaos, Solitons & Fractals 2004;20:995–1004].     

Hopf bifurcation in a predator-prey system with Holling type III functional response and time delays

This paper is concerned with a delayed predator-prey system with modified Leslie-Gower and Holling type III schemes. By analyzing the associated characteristic equation, its local stability and the existence of periodic solutions via Hopf bifurcation with respect to both delays are obtained. Based on the normal form method and center manifold theorem, the formulaes for determining the direction of Hopf bifurcation and the stability of bifurcating periodic solutions are derived. Finally, some numerical simulations to illustrate the theoretical analysis are also carried out.     

Dynamic behaviour of a delayed predator-prey model with harvesting

In this paper, we analyze the dynamics of a delayed predator-prey system in the presence of harvesting. This is a modified version of the Leslie-Gower and Holling-type II scheme. The main result is given in terms of local stability, global stability, influence of harvesting and bifurcation. Direction of Hopf bifurcation and the stability of bifurcating periodic solutions are also studied by using the normal form method and center manifold theorem.     

一类捕食者-食饵模型的稳定性及Hopf分支

研究一类具有时滞和阶段结构的捕食模型的稳定性和Hopf分支.以滞量为参数,得到了系统正平衡点的稳定性和Hopf分支存在的充分条件.利用规范型和中心流形定理,给出了确定Hopf分支方向和分支周期解的稳定性的计算公式.     

A stage-structured predator-prey system with time delay

A stage-structured predator-prey system with time delay is considered. By analyzing the corresponding characteristic equation, the local stability of a positive equilibrium is investigated. The existence of Hopf bifurcations is established. Formulae are derived to determine the direction of bifurcations and the stability of bifurcating periodic solutions by using the normal form theory and center manifold theorem. Numerical simulations are carried out to illustrate the theoretical results. Based on the global Hopf bifurcation theorem for general functional differential equations, the global existence of periodic solutions is established.     

A boundary value problem of thermal convection

The two-dimensional thermal convection of a viscous fluid in a rectangular region is analyzed using the Boussinesq theory, and the Rayleigh boundary conditions. The existence of a unique convention state bifurcating from each eigenvalue of the linearized theory is established by using the Morse lemma. This establishes the validity of the formal perturbation method for determining the convection states. The effects of imperfections on the transition from the conduction to the convection states are studied. A theorem of Thom is used to justify a previous asymptotic expansion method near the critical Rayleigh number.     

Hopf bifurcation and stability of periodic solutions for van der Pol equation with time delay

In this paper, the van der Pol equation with a time delay is considered, where the time delay is regarded as a parameter. It is found that Hopf bifurcation occurs when this delay passes through a sequence of critical value. A formula for determining the direction of the Hopf bifurcation and the stability of bifurcating periodic solutions is given by using the normal form method and center manifold theorem.     

Positive solutions bifurcating from zero solution in a Lotka-Volterra competitive system with cross-diffusion effects

A strongly coupled elliptic system under the homogeneous Dirichlet boundary condition denoting the steady-state system of the Lotka-Volterra two-species competitive system with cross-diffusion effects is considered. By using the implicit function theorem and the Lyapunov-Schmidt reduction method, the existence of the positive solutions bifurcating from the trivial solution is obtained. Furthermore, the stability of the bifurcating positive solutions is also investigated by analyzing the associated characteristic equation.     

Hopf bifurcation in a control system for the Washout filter-based delayed neural equation

Washout filter is a simple filter that can be designed easily. In this paper, a system for controlling a neural equation with discrete time delay based on Washout filter is presented. The transcendental equation of the corresponding linearized system is analyzed. In this control system, it is found that Hopf bifurcation occurs when the control parameters are chosen properly and that a chaotic orbit can be controlled to a stable periodic solution. The stability condition for bifurcating periodic solutions and the direction of Hopf bifurcation are studied by applying the normal form theory and the center manifold theorem. Some numerical results are also presented to illustrate the correctness of our results.