Modelling and analysis of a multiple delayed exploited ecosystem towards coexistence perspective

This paper describes a multiple delayed modified Leslie–Gower type predator–prey system with a strong Allee effect in prey population growth. Non-selective effort is used to harvest the population. The dynamical characteristics of the delay induced system are rigorously studied using mathematical tools. The existence of coexistence equilibria is ensured, and the dynamic behavior of the system is investigated around coexistence equilibria. Uniform strong persistence and permanence of the system are discussed in order to ensure long-term survival of the species. The stability of the delay preserved system is investigated. Sufficient conditions are derived for local and global stability of the system. The existence of Hopf bifurcation phenomenon is examined around interior equilibria of the system. Subsequently, we use normal form method and center manifold theorem to examine the nature of the Hopf bifurcation. Finally, numerical simulations are carried out to validate the analytical findings.     

Stability and Hopf bifurcation analysis for an energy resource system

This paper investigates the dynamical behaviors for a four-dimensional energy resource system with time delay, especially in terms of equilibria analyses and Hopf bifurcation analysis. By setting the time delay as a bifurcation parameter, it is shown that Hopf bifurcation would occur when the time delay exceeds a sequence of critical values. Furthermore, the stability and direction of the Hopf bifurcation are determined via the normal form theory and the center manifold reduction theorem. Numerical examples are given in the end of the paper to verify the theoretical results.     

Bifurcation and chaos of a delayed predator-prey model with dormancy of predators

In this paper, a delayed predator-prey model with dormancy of predators is investigated. It shows that time delay in the prey-species growth can lead to the occurrence of Hopf bifurcation with stability switches at a coexistence equilibrium. The computing formulas of stability and direction of the Hopf bifurcating periodic solutions are given. Under appropriate conditions, the uniform persistence of this model with time delay is proved. In this simple model, multiple periodic solutions coexist. Through numerical simulation, it is shown that different values of time delay can generate or eliminate chaos. Biologically, our results imply that dynamical behaviors of this system with time delay strongly depend on the initial density of this model and the time delay of the growth of the prey.     

Multiple scales and normal forms in a ring of delay coupled oscillators with application to chaotic Hindmarsh–Rose neurons

We study the appearance and stability of spatiotemporal periodic patterns like phase-locked oscillations, mirror-reflecting waves, standing waves, in-phase or antiphase oscillations, and coexistence of multiple patterns, in a ring of bidirectionally delay coupled oscillators. Hopf bifurcation, Hopf–Hopf bifurcation, and the equivariant Hopf bifurcation are studied in the viewpoint of normal forms obtained by using the method of multiple scales which is a kind of perturbation technique, thus a clear bifurcation scenario is depicted. We find time delay significantly affects the dynamics and induces rich spatiotemporal patterns. With the help of the unfolding system near Hopf–Hopf bifurcation, it is confirmed in some regions two kinds of stable oscillations may coexist. These phenomena are shown for the delay coupled limit cycle oscillators as well as for the delay coupled chaotic Hindmarsh–Rose neurons.     

Bifurcation analysis of 4-axle rail vehicle models in a curved track

The dynamics of a diffusive predator–prey model with time delay and Michaelis–Menten-type harvesting subject to Neumann boundary condition is considered. Turing instability and Hopf bifurcation at positive equilibrium for the system without delay are investigated. Time delay-induced instability and Hopf bifurcation are also discussed. By the theory of normal form and center manifold, conditions for determining the bifurcation direction and the stability of bifurcating periodic solution are derived. Some numerical simulations are carried out for illustrating the theoretical results.     

Dynamics analysis of a non-smooth Filippov pest-natural enemy system with time delay

In this paper, we propose a non-smooth Filippov system that describes the interaction of the pest and natural enemy with considering time delay, which represents the change in the growth rate of natural enemies before it is released to prey on pests. When the number of the pest is below the threshold, no control is applied; otherwise, control measures will be adopted. We discuss the stability of the equilibria and the existence of Hopf bifurcation. The results show that the Hopf bifurcation occurs when the time delay passes through some critical values. By applying the Filippov convex method, we obtain the dynamics of the sliding mode. The solutions of the system eventually tend toward the regular equilibrium, the pseudo-equilibrium or a standard periodic solution. Numerical simulations show that time delay plays an important role in local and global sliding bifurcations. We can obtain boundary focus bifurcations from boundary node bifurcations by varying time delay. Furthermore, touching, buckling and crossing bifurcations can be obtained frequently by increasing time delay. The results can provide some insights in pest control.

     

Dynamics in two nonsmooth predator–prey models with threshold harvesting

Considering a good pest control program should reduce the pest to levels acceptable to the public, we investigate the threshold harvesting policy on pests in two predator–prey models. Both models are nonsmooth and the aim of this paper is to provide how threshold harvesting affects the dynamics of the two systems. When the harvesting threshold is larger than some positive level, the harvesting does not affect the ecosystem; when the harvesting threshold is less than the level, the model has complex dynamics with multiple coexistence equilibria, limit cycle, bistability, homoclinic orbit, saddle-node bifurcation, transcritical bifurcation, subcritical and supercritical Hopf bifurcation, Bogdanov–Takens bifurcation, and discontinuous Hopf bifurcation. Firstly, we provide the complete stability analysis and bifurcation analysis for the two models. Furthermore, some numerical simulations are given to illustrate our results. Finally, it is found that harvesting lowers the level of both species for natural enemy–pest system while raises the densities of both species for the pest–crop system. It is seen that the threshold harvesting policy of the enemy system is more effective than the crop system.     

Stability and bifurcation in a generalized delay prey–predator model

The present paper considers a generalized prey–predator model with time delay. It studies the stability of the nontrivial positive equilibrium and the existence of Hopf bifurcation for this system by choosing delay as a bifurcation parameter and analyzes the associated characteristic equation. The researcher investigates the direction of this bifurcation by using an explicit algorithm. Eventually, some numerical simulations are carried out to support the analytical results.     

Optimal harvesting and complex dynamics in a delayed eco-epidemiological model with weak Allee effects

In this article, an eco-epidemiological system with weak Allee effect and harvesting in prey population is discussed by a system of delay differential equations. The delay parameter regarding the time lag corresponds to the predator gestation period. Mathematical features such as uniform persistence, permanence, stability, Hopf bifurcation at the interior equilibrium point of the system is analyzed and verified by numerical simulations. Bistability between different equilibrium points is properly discussed. The chaotic behaviors of the system are recognized through bifurcation diagram, Poincare section and maximum Lyapunov exponent. Our simulation results suggest that for increasing the delay parameter, the system undergoes chaotic oscillation via period doubling. We also observe a quasi-periodicity route to chaos and complex dynamics with respect to Allee parameter; such behavior can be subdued by the strength of the Allee effect and harvesting effort through period-halving bifurcation. To find out the optimal harvesting policy for the time delay model, we consider the profit earned by harvesting of both the prey populations. The effect of Allee and gestation delay on optimal harvesting policy is also discussed.     

Rich dynamics in a non-local population model over three patches

We consider a system of nonlinear delay differential equations that describes the growth of the mature population of a species with age-structure living over three patches. We analyze existence of non-negative homogeneous equilibria and their stability and discuss possible Hopf bifurcation from these equilibria. More precisely, by employing both the standard Hopf bifurcation theory and the symmetric bifurcation theory for functional differential equations, we obtain very rich dynamics for the system, including bistable equilibria, transient oscillations, synchronous periodic solutions, phase-locked periodic solutions, mirror-reflecting waves and standing waves.     

A delayed ratio-dependent predator–prey model of interacting populations with Holling type III functional response

In this article, we study a ratio-dependent predator–prey model described by a Holling type III functional response with time delay incorporated into the resource limitation of the prey logistic equation. This investigation includes the influence of intra-species competition among the predator species. All the equilibria are characterized. Qualitative behavior of the complicated singular point (0,0) in the interior of the first quadrant is investigated by means of a blow-up transformation. Uniform persistence, stability, and Hopf bifurcation at the positive equilibrium point of the system are examined. Global asymptotic stability analyses of the positive equilibrium point by the Bendixon–Dulac criterion for non-delayed model and by constructing a suitable Lyapunov functional for the delayed model are carried out separately. We perform a numerical simulation to validate the applicability of the proposed mathematical model and our analytical findings.     

Stability and Hopf bifurcation in a three-neuron unidirectional ring with distributed delays

In this paper, we consider the effect of distributed delays in a three-neuron unidirectional ring. Sufficient conditions for existence of unique equilibrium, multiple equilibria and their local stability are derived. Taking the average delay as a bifurcation parameter, we find two critical values at which the system undergoes Hopf bifurcations. The orbital asymptotic stability of the Hopf bifurcating periodic solutions is investigated by using the method of multiple scales. The global Hopf bifurcation is also studied. Finally, the theoretical results are illustrated by some numerical simulations.     

Bifurcation of a predator–prey model with disease in the prey

In this paper, a predator–prey model with disease in the prey is considered. Assume that the predator eats only the infected prey, and the incidence rate is nonlinear. We study the dynamics of the model in terms of local analysis of equilibria and bifurcation analysis of a boundary equilibrium and a positive equilibrium. We discuss the Bogdanov–Takens bifurcation near the boundary equilibrium and the Hopf bifurcation near the positive equilibrium; numerical simulation results are given to support the theoretical predictions.     

van der Pol型时滞系统的两参数余维一Hopf分岔及其稳定性

研究具有三次非线性时滞项的ｖａｎ ｄｅｒ Ｐｏｌ型时滞系统随两参数（时滞量和增益系数）余维一Ｈｏｐｆ分岔,说明了线性化特性方程随两参数变化时的根的分布和Ｈｏｐｆ分岔存在性;通过构造中心流形并且使用范式方法确定出Ｈｏｐｆ分岔的方向以及周期解的稳定性;分析了时滞量对所论系统发生Ｈｏｐｆ分岔的影响。     

Stochastic Hopf bifurcation of quasi-integrable Hamiltonian systems with multi-time-delayed feedback control and wide-band noise excitations

The stochastic Hopf bifurcation of multi-degree-of-freedom (MDOF) quasi-integrable Hamiltonian systems with multi-time-delayed feedback control subject to wide-band noise excitations is studied. First, the time-delayed feedback control forces are expressed approximately in terms of the system state variables without time delay and the system is converted into an ordinary quasi-integrable Hamiltonian system. The averaged It? stochastic differential equations are derived by using the stochastic averaging method for quasi-integrable Hamiltonian systems. Then the expression for average bifurcation parameter of the averaged system is obtained approximately and a criterion for determining the stochastic Hopf bifurcation induced by time-delayed feedback control forces in the original system using average bifurcation parameter is proposed. An example is worked out in detail to illustrate the criterion and its validity and to show the effect of time delay in feedback control on stochastic Hopf bifurcation of the system.     

Effort dynamics of a delay-induced prey–predator system with reserve

This paper describes a prey?Cpredator fishery system with prey dispersal in a two-patch environment, one of which is a free fishing zone and the other a protected zone. The proposed system reflects the dynamic interaction between the net economic revenue and the fishing effort used to harvest the population in presence of a suitable tax. Local as well as global stability of the system is analyzed. The optimal taxation policy is formulated and solved with the help of Pontryagin??s maximal principle. The objective of the paper is to achieve the sustainability of the fishery, keeping the ecological balance, and maximize the monetary social benefit. The dynamical behavior of the delay system is further analyzed through incorporating discrete type gestational delay of predators, and the existence of Hopf bifurcation phenomenon is checked at the interior equilibrium point. Moreover, we use normal form method and center manifold theorem to examine the nature of the Hopf bifurcation. Theoretical results are verified with the help of numerical examples and graphical illustrations.     

One-to-three resonant Hopf bifurcations of a maglev system

This paper studies the dynamics of a maglev system around 1:3 resonant Hopf–Hopf bifurcations. When two pairs of purely imaginary roots exist for the corresponding characteristic equation, the maglev system has an interaction of Hopf–Hopf bifurcations at the intersection of two bifurcation curves in the feedback control parameter and time delay space. The method of multiple time scales is employed to drive the bifurcation equations for the maglev system by expressing complex amplitudes in a combined polar-Cartesian representation. The dynamics behavior in the vicinity of 1:3 resonant Hopf–Hopf bifurcations is studied in terms of the controller’s parameters (time delay and two feedback control gains). Finally, numerical simulations are presented to support the analytical results and demonstrate some interesting phenomena for the maglev system.     

Delayed feedback control and bifurcation analysis of Rossler chaotic system

In this paper, from the view of stability and chaos control, we investigate the Rossler chaotic system with delayed feedback. At first, we consider the stability of one of the fixed points, verifying that Hopf bifurcation occurs as delay crosses some critical values. Then, for determining the stability and direction of Hopf bifurcation we derive explicit formulae by using the normal-form theory and center manifold theorem. By designing appropriate feedback strength and delay, one of the unstable equilibria of the Rossler chaotic system can be controlled to be stable, or stable bifurcating periodic solutions occur at the neighborhood of the equilibrium. Finally, some numerical simulations are carried out to support the analytic results.     

Resonant double Hopf bifurcation in a diffusive Ginzburg–Landau model with delayed feedback

We investigate the resonant double Hopf bifurcation in a diffusive complex Ginzburg–Landau model with delayed feedback and phase shift. The conditions for the existence of resonant double Hopf bifurcation are obtained by analyzing the roots’ distribution of the characteristic equation, and a general formula to determine the bifurcation point is given. For the cases of 1:2 and 1:3 resonance, we choose time delay, feedback strength and phase shift as bifurcation parameters and derive the normal forms which are proved to be the same as those in non-resonant cases. The impact of cubic terms on the unfolding types is discussed after obtaining the normal form till 3rd order. By fixing phase shift, we find that varying time delay and feedback strength simultaneously can induce the coexistence of two different periodic solutions, the existence of quasi-periodic solutions and strange attractors. Also, the effects on the existence of transient quasi-periodic solution exerted by the phase shift are illustrated.

     

Stability and bifurcation analysis of delay coupled Van der Pol–Duffing oscillators

In this paper, we aim to investigate the dynamics of a system of Van der Pol–Duffing oscillators with delay coupling. First, taking the time delay as a bifurcation parameter, the stability of the equilibrium, and the existence of Hopf bifurcation are investigated. Then using the center manifold reduction technique and normal form theory, we give the direction of the Hopf bifurcation. And then by means of the symmetric bifurcation theory for delay differential equations and the representation theory of groups, we claim the bifurcation periodic solution induced by time delay is antiphase locked oscillation. Finally, at the end of the paper, numerical simulations are carried out to support our theoretical analysis.