Die Phlorglucide von Dryopteris viliarii (Bell.) Woynar und anderer Farne der Gattung Dryopteris sowie die mögliche Abstammung von D. filix-mas (L.) Schott

The phloroglucides of the ferns of the Dryopteris filix-mas complex; i.e. D. abbreviata, D. filix-mas s. str. and D. borreri (diploid and triploid) as well as those of D. villarii, subspvillarii and subsp. pallida were reinvestigated with improved semiquantitative analytical methods. The phloroglucides of D. aitoniana and D. athamantica were examined for the first time. The results (table 1) are compatible with the hypothesis, without proving it, that the allotetraploid species D. filix-mas s. str. originated from a hybrid of D. abbreviata with D. villard (with subsequent doubling of its chromosomes) and the apogamous triploid D. remota from D. assimils with diploid D. borreri, and the equally apogamous triploid D. borreri from a hybrid of D. abbreviata (or a related diploid sexual taxon) with diploid D. borreri. D. aitoniana contains a large amount of trisflavaspidic acid but no filixic acid and differs in this respect from the three representatives of the D. filix-mas complex.