一类三维神经元模型的分支研究

考虑一类三维神经元模型的分支问题.利用常微分方程的定性与分支理论的知识,讨论了模型的平衡点个数及其稳定性,主要分析了平衡点的Hopf分支和Bogdanov-Takens分支,并得到了相应的鞍结点分支曲线,Hopf分支曲线与同宿分支曲线.     

Slow-fast Bogdanov-Takens bifurcations

In this paper we study perturbations from planar vector fields having a line of zeros and representing a singular limit of Bogdanov-Takens (BT) bifurcations. We introduce, among other precise definitions, the notion of slow-fast BT-bifurcation and we provide a complete study of the bifurcation diagram and the related phase portraits. Based on geometric singular perturbation theory, including blow-up, we get results that are valid on a uniform neighborhood both in parameter space and in the phase plane.     

Analyses of Bifurcations and Stability in a Predator-prey System with Holling Type-Ⅳ Functional Response

In this paper the dynamical behaviors of a predator-prey system with Holling Type-Ⅳfunctionalresponse are investigated in detail by using the analyses of qualitative method,bifurcation theory,and numericalsimulation.The qualitative analyses and numerical simulation for the model indicate that it has a unique stablelimit cycle.The bifurcation analyses of the system exhibit static and dynamical bifurcations including saddle-node bifurcation,Hopf bifurcation,homoclinic bifurcation and bifurcation of cusp-type with codimension two(ie,the Bogdanov-Takens bifurcation),and we show the existence of codimension three degenerated equilibriumand the existence of homoclinic orbit by using numerical simulation.     

Dynamical behavior of an epidemic model with a nonlinear incidence rate

In this paper, we study the global dynamics of an epidemic model with vital dynamics and nonlinear incidence rate of saturated mass action. By carrying out global qualitative and bifurcation analyses, it is shown that either the number of infective individuals tends to zero as time evolves or there is a region such that the disease will be persistent if the initial position lies in the region and the disease will disappear if the initial position lies outside this region. When such a region exists, it is shown that the model undergoes a Bogdanov-Takens bifurcation, i.e., it exhibits a saddle-node bifurcation, Hopf bifurcations, and a homoclinic bifurcation. Existence of none, one or two limit cycles is also discussed.     

Bifurcations of a Leslie-Gower prey-predator system with ratio-dependent Holling IV functional response and prey harvesting

The dynamics of a Leslie-Gower prey-predator system with ratio-dependent Holling IV functional response and constant harvesting rate of prey are taken into account. The results developed in this article reveal far richer dynamics compared with the system without harvesting. We first make qualitative and bifurcation analysis of the system without harvesting and show that the system has a weak focus of multiplicity at most 2, at which a Hopf bifurcation occurs. However, the system with harvesting has four nonhyperbolic equilibria for some parameter values, such as two saddle-node, a cusp, and a weak focus of multiplicity at most 4, and exhibits two saddle-node bifurcations, a Bogdanov-Takens bifurcation of codimension 2, and a Hopf bifurcation. It reveals that there exist some critical harvesting values such that the species are in danger of extinction when the harvesting rate is greater than the critical values, which indicates that the dynamics of the system are sensitive to the constant prey harvesting. Moreover, numerical simulations are presented to illustrate our theoretical results.     

Modeling and analysis of recurrent autoimmune disease

In this paper, we consider dynamics and bifurcations in two HIV models with cell-to-cell interaction. The difference between the two models lies in the inclusion or omission of the effect of involvement. Particular attention is focused on the effects due to the cell-to-cell transmission and the effect of the involvement. We investigate the local and global stability of equilibria of the two models and give a comparison. We derive the existence condition for Hopf bifurcation and prove no Bogdanov-Takens bifurcation in this system. In particular, we show that the system exhibits the recurrence phenomenon, yielding complex dynamical behavior. It is also shown that the effect of the involvement is the main cause of the periodic symptoms in HIV or malaria disease. Moreover, it is shown that the increase of cell-to-cell interaction may be the main factor causing Hopf bifurcation to disappear, and thus eliminating oscillation behavior. Finally, numerical simulations are present to demonstrate our theoretical results.     

Center manifolds theorem for parameterized delay differential equations with applications to zero singularities

The goal of this paper is to develop a center manifold theory for delay differential equations with parameters. As applications, we use the center manifold theorem to establish fold and Bogdanov-Takens bifurcations. In particular, we obtain the versal unfoldings of delayed predator-prey systems with predator harvesting at the Bogdanov-Takens singularity.     

Bifurcations and Chaos in a Discrete Predator-prey System with Holling Type-IV Functional Response

A discrete predator-prey system with Holling type-IV functional response obtained by the Euler method is first investigated. The conditions of existence for fold bifurcation, flip bifurcation and Hopf bifurcation are derived by using the center manifold theorem and bifurcation theory. Furthermore, we give the condition for the occurrence of codimension-two bifurcation called the Bogdanov-Takens bifurcation for fixed points and present approximate expressions for saddle-node, Hopfand homoclinic bifurcation sets near the Bogdanov-Takens bifurcation point. We also show the existence of degenerated fixed point with codimension three at least. The numerical simulations, including bifurcation diagrams, phase portraits, and computation of maximum Lyapunov exponents, not only show the consistence with the theoretical analysis but also exhibit the rich and complex dynamical behaviors such as the attracting invariant circle, period-doubling bifurcation from period-2,3,4 orbits.interior crisis, intermittency mechanic, and sudden disappearance of chaotic dynamic.     

Dynamics of SIR epidemic models with horizontal and vertical transmissions and constant treatment rates

We investigate the dynamics and bifurcations of SIR epidemic model with horizontal and vertical transmissions and constant treatment rates. It is proved that such SIR epidemic model have up to two positive epidemic equilibria and has no positive disease-free equilibria. We find all the ranges of the parameters involved in the model under which the equilibria of the model are positive. By using the qualitative theory of planar systems and the normal form theory, the phase portraits of each equilibria are obtained. We show that the equilibria of the epidemic system can be saddles, stable nodes, stable or unstable focuses, weak centers or cusps. We prove that the system has the Bogdanov-Takens bifurcations, which exhibit saddle-node bifurcations, Hopf bifurcations and homoclinic bifurcations.     

Bogdanov-Takens bifurcation in a delayed Michaelis-Menten type ratio-dependent predator-prey system with prey harvesting

In this paper, we study a delayed Michaelis-Menten Type ratio-dependent predator-prey model with prey harvesting. By considering the characteristic equation associated with the nonhyperbolic equilibrium, the critical value of the parameters for the Bogdanov-Takens bifurcation is obtained. The conditions for the characteristic equation having negative real parts are discussed. Using the normal form theory of Bogdanov-Takens bifurcation for retarded functional differential equations, the corresponding normal form restricted to the associated two-dimensional center manifold is calculated and the versal unfolding is considered. The parameter conditions for saddle-node bifurcation, Hopf bifurcation and homoclinic bifurcation are obtained. Numerical simulations are given to support the analytical results.     

Dynamics in a ratio-dependent predator-prey model with predator harvesting

The objective of this paper is to study systematically the dynamical properties of a ratio-dependent predator-prey model with nonzero constant rate predator harvesting. It is shown that the model has at most two equilibria in the first quadrant and can exhibit numerous kinds of bifurcation phenomena, including the bifurcation of cusp type of codimension 2 (i.e., Bogdanov-Takens bifurcation), the subcritical and supercritical Hopf bifurcations. These results reveal far richer dynamics compared to the model with no harvesting and different dynamics compared to the model with nonzero constant rate prey harvesting in [D. Xiao, L. Jennings, Bifurcations of a ratio-dependent predator-prey system with constant rate harvesting, SIAM Appl. Math. 65 (2005) 737-753]. Biologically, it is shown that nonzero constant rate predator harvesting can prevent mutual extinction as a possible outcome of the predator prey interaction, and remove the singularity of the origin, which was regarded as “pathological behavior” for a ratio-dependent predator prey model in [P. Yodzis, Predator-prey theory and management of multispecies fisheries, Ecological Applications 4 (2004) 51-58].     

一类具有非线性发生率和治疗函数的传染病模型研究

传染病动力学系统的数学建模中,合理的使用非线性发生率往往更能使模型与实际相吻合.并且在实际的疾病防治过程中,由于受到空间人力物力资源的影响一般存在最大治疗容量的限制.结合这两种情况建立了一类含非线性发生率和最大治疗容量限制的传染病模型.通过分析这个模型,得到无病平衡点和正平衡点的存在性、稳定性.进一步取发生率和治疗系统达到最大容量时的感染者人数作为分支参数,得到了Hopf分支和Bogdanov-Takens分支的存在条件,并进行了数值模拟.     

Two-parameter bifurcations in a network of two neurons with multiple delays

We consider a network of two coupled neurons with delayed feedback. We show that the connection topology of the network plays a fundamental role in classifying the rich dynamics and bifurcation phenomena. Regarding eigenvalues of the connection matrix as bifurcation parameters, we obtain codimension 1 bifurcations (including a fold bifurcation and a Hopf bifurcation) and codimension 2 bifurcations (including fold-Hopf bifurcations and Hopf-Hopf bifurcations). We also give concrete formulae for the normal form coefficients derived via the center manifold reduction that give detailed information about the bifurcation and stability of various bifurcated solutions. In particular, we obtain stable or unstable equilibria, periodic solutions, quasi-periodic solutions, and sphere-like surfaces of solutions. We also show how to evaluate critical normal form coefficients from the original system of delay-differential equations without computing the corresponding center manifolds.     

Bifurcation of an eco-epidemiological model with a nonlinear incidence rate

A predator-prey system with disease in the prey is considered. Assume that the incidence rate is nonlinear, we analyse the boundedness of solutions and local stability of equilibria, by using bifurcation methods and techniques, we study Bogdanov-Takens bifurcation near a boundary equilibrium, and obtain a saddle-node bifurcation curve, a Hopf bifurcation curve and a homoclinic bifurcation curve. The Hopf bifurcation and generalized Hopf bifurcation near the positive equilibrium is analyzed, one or two limit cycles is also discussed.     

Periodic orbits and chaos in fast-slow systems with Bogdanov-Takens type fold points

The existence of stable periodic orbits and chaotic invariant sets of singularly perturbed problems of fast-slow type having Bogdanov-Takens bifurcation points in its fast subsystem is proved by means of the geometric singular perturbation method and the blow-up method. In particular, the blow-up method is effectively used for analyzing the flow near the Bogdanov-Takens type fold point in order to show that a slow manifold near the fold point is extended along the Boutroux's tritronquée solution of the first Painlevé equation in the blow-up space.     

Bifurcation from periodic solutions with spatiotemporal symmetry, including resonances and mode interactions

We study local bifurcation in equivariant dynamical systems from periodic solutions with a mixture of spatial and spatiotemporal symmetries.In previous work, we focused primarily on codimension one bifurcations. In this paper, we show that the techniques used in the codimension one analysis can be extended to understand also higher codimension bifurcations, including resonant bifurcations and mode interactions. In particular, we present a general reduction scheme by which we relate bifurcations from periodic solutions to bifurcations from fixed points of twisted equivariant diffeomorphisms, which in turn are linked via normal form theory to bifurcations from equilibria of equivariant vector fields.We also obtain a general theory for bifurcation from relative periodic solutions and we show how to incorporate time-reversal symmetries into our framework.     

BIFURCATIONS OF DEGENERATE SINGULAR POINT OF A PREDATOR-PREY SYSTEM WITH HOLLING-IV FUNCTIONAL RESPONSE

A predator-prey system with Holling-IV functional response is investigated. It is shown that the system has a positive equilibrium?which is a cusp of co-dimension 2 under certain conditions. When the parameters vary in a small neighborhood of the values of parameters, the model undergoes the Bogdanov-Takens bifurcation. Different kinds of bifurcation phenomena are exhibited, which include the saddle~node bifurcation, the Hopf bifurcation and the homo-clinic bifurcation. Some computer simulations are presented to illustrate the conclusions.     

Bifurcation analysis of an epidemic model with nonlinear incidence

In this paper, we consider an epidemic model with the nonlinear incidence of a sigmoidal function. By mathematical analysis, it is shown that the model exhibits the bistability and undergoes the Hopf bifurcation and the Bogdanov-Takens bifurcation. By numerical simulations, it is found that the incidence rate can induce multiple limit cycles, and a little change of the parameter could lead to quite different bifurcation structures.     

Synchronous dynamics of two coupled oscillators with inhibitory-to-inhibitory connection

We investigate the behaviour of a neural network model consisting of two coupled oscillators with delays and inhibitory-to-inhibitory connections. We consider the absolute synchronization and show that the connection topology of the network plays a fundamental role in classifying the rich dynamics and bifurcation phenomena. Regarding eigenvalues of the connection matrix as bifurcation parameters, we obtain codimension one bifurcations (including fold bifurcation and Hopf bifurcation) and codimension two bifurcation (including fold-Hopf bifurcations and Hopf–Hopf bifurcations). Based on the normal form theory and center manifold reduction, we obtain detailed information about the bifurcation direction and stability of various bifurcated equilibria as well as periodic solutions with some kinds of spatio-temporal patterns. Numerical simulation is also given to support the obtained results.     

Bogdanov-Takens bifurcation in a Leslie-Gower predator-prey model with prey harvesting

This paper discuss the cusp bifurcation of codimension 2 （i.e. Bogdanov-Takens bifurcation） in a Leslie~Gower predator-prey model with prey harvesting, which was not revealed by Zhu and Lan [Phase portraits, Hopf bifurcation and limit cycles of Leslie-Gower predator-prey systems with harvesting rates, Discrete and Continuous Dynamical Systems Series B. 14（1） （2010）, 289-306]. It is shown that there are different parameter values for which the model has a limit cycle or a homoclinic loop.