Bifurcation analysis of a generalized Gause model with prey harvesting and a generalized Holling response function of type III

In this paper we study a generalized Gause model with prey harvesting and a generalized Holling response function of type III: . The goal of our study is to give the bifurcation diagram of the model. For this we need to study saddle-node bifurcations, Hopf bifurcation of codimension 1 and 2, heteroclinic bifurcation, and nilpotent saddle bifurcation of codimension 2 and 3. The nilpotent saddle of codimension 3 is the organizing center for the bifurcation diagram. The Hopf bifurcation is studied by means of a generalized Liénard system, and for b=0 we discuss the potential integrability of the system. The nilpotent point of multiplicity 3 occurs with an invariant line and can have a codimension up to 4. But because it occurs with an invariant line, the effective highest codimension is 3. We develop normal forms (in which the invariant line is preserved) for studying of the nilpotent saddle bifurcation. For b=0, the reversibility of the nilpotent saddle is discussed. We study the type of the heteroclinic loop and its cyclicity. The phase portraits of the bifurcations diagram (partially conjectured via the results obtained) allow us to give a biological interpretation of the behavior of the two species.     

Bifurcations of a Leslie-Gower prey-predator system with ratio-dependent Holling IV functional response and prey harvesting

The dynamics of a Leslie-Gower prey-predator system with ratio-dependent Holling IV functional response and constant harvesting rate of prey are taken into account. The results developed in this article reveal far richer dynamics compared with the system without harvesting. We first make qualitative and bifurcation analysis of the system without harvesting and show that the system has a weak focus of multiplicity at most 2, at which a Hopf bifurcation occurs. However, the system with harvesting has four nonhyperbolic equilibria for some parameter values, such as two saddle-node, a cusp, and a weak focus of multiplicity at most 4, and exhibits two saddle-node bifurcations, a Bogdanov-Takens bifurcation of codimension 2, and a Hopf bifurcation. It reveals that there exist some critical harvesting values such that the species are in danger of extinction when the harvesting rate is greater than the critical values, which indicates that the dynamics of the system are sensitive to the constant prey harvesting. Moreover, numerical simulations are presented to illustrate our theoretical results.     

Hopf bifurcation in a diffusive predator-prey model with herd behavior and prey harvesting

In this paper, the dynamics of a diffusive delayed predator-prey model with herd behavior and prey harvesting subject to the homogeneous Neumann boundary condition is considered. Firstly, choosing the harvesting term as a bifurcation parameter, then we obtain the existence and the stability of the equilibrium by analyzing the distribution of the roots of associated characteristic equation. Secondly, time delay is regarding as a bifurcation parameter, and the use of the normal form theory and center manifold theorem, the existence, stability and direction of bifurcating periodic solutions are all demonstrated detailly. Finally, summarizing some numerical simulations to illustrate the theoretical analysis.     

Bogdanov-Takens bifurcation in a delayed Michaelis-Menten type ratio-dependent predator-prey system with prey harvesting

In this paper, we study a delayed Michaelis-Menten Type ratio-dependent predator-prey model with prey harvesting. By considering the characteristic equation associated with the nonhyperbolic equilibrium, the critical value of the parameters for the Bogdanov-Takens bifurcation is obtained. The conditions for the characteristic equation having negative real parts are discussed. Using the normal form theory of Bogdanov-Takens bifurcation for retarded functional differential equations, the corresponding normal form restricted to the associated two-dimensional center manifold is calculated and the versal unfolding is considered. The parameter conditions for saddle-node bifurcation, Hopf bifurcation and homoclinic bifurcation are obtained. Numerical simulations are given to support the analytical results.     

Hopf bifurcation analysis for a delayed predator-prey system with a prey refuge and selective harvesting

In this paper, a delayed predator-prey system with Holling type III functional response incorporating a prey refuge and selective harvesting is considered. By analyzing the corresponding characteristic equations, the conditions for the local stability and existence of Hopf bifurcation for the system are obtained, respectively. By utilizing normal form method and center manifold theorem, the explicit formulas which determine the direction of Hopf bifurcation and the stability of bifurcating period solutions are derived. Finally, numerical simulations supporting the theoretical analysis are given.     

Zero‐Hopf bifurcation in the Volterra‐Gause system of predator‐prey type

We prove that the Volterra‐Gause system of predator‐prey type exhibits 2 kinds of zero‐Hopf bifurcations for convenient values of their parameters. In the first, 1 periodic solution bifurcates from a zero‐Hopf equilibrium, and in the second, 4 periodic solutions bifurcate from another zero‐Hopf equilibrium. This study is done using the averaging theory of second order.     

Hopf bifurcation analysis of a predator-prey system with Holling type IV functional response and time delay

This paper is concerned with a predator-prey system with Holling type IV functional response and time delay. Our aim is to investigate how the time delay affects the dynamics of the predator-prey system. By choosing the delay as a bifurcation parameter, the local asymptotic stability of the positive equilibrium and existence of local Hopf bifurcations are analyzed. Based on the normal form and the center manifold theory, the formulaes for determining the properties of Hopf bifurcation of the predator-prey system are derived. Finally, to support these theoretical results, some numerical simulations are given to illustrate the results.     

Dynamics of a delayed predator-prey model with Allee effect and Holling type II functional response

In this paper, a delayed with Holling type II functional response (Beddington-DeAngelis) and Allee effect predator-prey model is considered. The growth of the prey is affected by the parameter $M$, which defines the Allee effect. In addition, the delay $\tau$ also influences the logistic growth of the prey, which can be interpreted as the maturity time or the gestation period. In the study of the characteristic equation, we observe that the delay $\tau$ also depends on the parameter $M$, which affects the dynamics in the prey population. Considering the delay as a bifurcation parameter, the local asymptotic stability of the positive equilibrium is investigated. On the other hand, we find that the system can also suffer a Hopf bifurcation in the positive equilibrium when the delay passes through a sequence of critical values. In particular, we study the direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions, an explicit algorithm is provided applying the normal form theory and center manifold reduction for the functional differential equations. Finally, numerical simulations that support the theoretical analysis are included.     

Melnikov function and limit cycle bifurcation from a nilpotent center

We investigate a general near-Hamiltonian system on the plane whose unperturbed system has a nilpotent center. Our main purpose is to give an algorithm for calculating the first coefficients of the expansion of the first order Melnikov function. We also give an application by using the method and obtain the number of limit cycles of a cubic system.     

Uniform persistence of multispecies ecological competition predator-pray system with Holling III type functional response

The main purpose of this paper is to study the persistence of the general multispecies competition predator-pray system with Holling Ⅲ type functional response. In this system, the competition among predator species and among prey species are simultaneously considered. By using the comparison theory and qualitative analysis, the sufficient conditions for uniform strong persistence are obtained.     

Turing instability and Hopf bifurcation in a diffusive Leslie–Gower predator–prey model

In this paper, a reaction‐diffusion predator–prey system that incorporates the Holling‐type II and a modified Leslie‐Gower functional responses is considered. For ODE, the local stability of the positive equilibrium is investigated and the specific conditions are obtained. For partial differential equation, we consider the dissipation and persistence of solutions, the Turing instability of the equilibrium solutions, and the Hopf bifurcation. By calculating the normal form, we derive the formulae, which can determine the direction and the stability of Hopf bifurcation according to the original parameters of the system. We also use some numerical simulations to illustrate our theoretical results. Copyright © 2016 John Wiley & Sons, Ltd.     

Multiplicity of periodic solutions for a delayed ratio-dependent predator-prey model with Holling type III functional response and harvesting terms

By using the generalized continuation theorem, the existence of four positive periodic solutions for a delayed ratio-dependent predator-prey model with Holling type III functional response

     

Bifurcation behaviours in a delayed three-species food-chain model with Holling type-II functional response

This article is concerned with the local stability of a positive equilibrium and the Hopf bifurcation of a delayed three-species food-chain system with the Holling type-II functional response. Some new sufficient conditions ensuring the local stability of a positive equilibrium and the existence of Hopf bifurcation for the system are established. Some explicit formulae are derived to determine the direction of bifurcations and the stability of bifurcating periodic solutions using the normal form theory and the centre manifold theory. Numerical simulations for justifying the theoretical analysis are also provided. Finally, biological explanations and main conclusions are included.     

Stability and Hopf bifurcation of a diffusive predator‐prey model with hyperbolic mortality

The dynamics of a reaction‐diffusion predator‐prey model with hyperbolic mortality and Holling type II response effect is considered. The stability of the positive equilibrium and the existence of Hopf bifurcation are investigated by analyzing the distribution of eigenvalues without diffusion. We also study the spatially homogeneous and nonhomogeneous periodic solutions through all parameters of the system which are spatially homogeneous. To verify our theoretical results, some numerical simulations are also presented. © 2015 Wiley Periodicals, Inc. Complexity 21: 34–43, 2016     

Global stability and Hopf bifurcation of a delayed eco‐epidemiological model with Holling type II functional response

In this paper, a delayed eco‐epidemiological model with Holling type II functional response is investigated. By analyzing corresponding characteristic equations, the local stability of each of the feasible equilibria and the existence of Hopf bifurcations at the disease‐free equilibrium, the susceptible predator‐free equilibrium and the endemic‐coexistence equilibrium are established, respectively. By means of Lyapunov functionals and LaSalle's invariance principle, sufficient conditions are derived for the global stability of the endemic‐coexistence equilibrium, the disease‐free equilibrium, the susceptible predator‐free equilibrium and the predator‐extinction equilibrium of the system, respectively. Numerical simulations are carried out to illustrate the theoretical results. Copyright © 2014 John Wiley & Sons, Ltd.     

Dynamics of a modified Leslie-Gower model with gestation effect and nonlinear harvesting

This study focuses on the dynamics of a modified Leslie-Gower predator-prey model where the intake rate of prey is by per capita predator according to Crowley-Martin functional response and prey is harvested through nonlinear harvesting strategy. Further the time-delay $(\tau)$ is imposed to utilize gestation period of predations. We investigate the permanence analysis of proposed system. The local stability of non-delayed model at all possible equilibrium points is studied. It is shown that the given model undergoes Hopf bifurcation around positive equilibrium point with respect to delay parameter $\tau$. Subsequently the stability of Hopf bifurcation and its direction are explored through normal and center manifold theories. The derived theoretical results are justified with the help of numerical simulations.     

Bifurcation analysis of a predator–prey model with memory-based diffusion

In this paper, we investigate the predator–prey model equipped with Fickian diffusion and memory-based diffusion of predators. The stability and bifurcation analysis explores the impacts of the memory-based diffusion and the averaged memory period on the dynamics near the positive steady state. Specifically, when the memory-based diffusion coefficient is less than a critical value, we show that the stability of the positive steady state can be destabilized as the average memory period increases, which leads to the occurrence of Hopf bifurcations. Moreover, we also analyze the bifurcation properties using the central manifold theorem and normal form theory. This allows us to prove the existence of stable spatially inhomogeneous periodic solutions arising from Hopf bifurcation. In addition, the sufficient and necessary conditions for the occurrence of stability switches are also provided.     

Ecological complexity and feedback control in a prey–predator system with Holling type III functional response

This article deals with a bioeconomic model of prey–predator system with Holling type III functional response. The dynamical behavior of the system is extensively discussed. Continuous type gestational delay of predators is incorporated in the system to study delay induced instability. It is observed that the system undergoes singularity induced bifurcation at interior equilibrium point when net economic revenue of the system increases through zero. State feedback controller is designed to stabilize the system at positive economic profit. Time delay is considered as a bifurcation parameter to prove the occurrence of Hopf bifurcation phenomenon in the neighborhood of the coexisting equilibrium point. Finally, some numerical simulations are carried out to verify the analytical results and the system is analyzed through graphical illustrations. © 2015 Wiley Periodicals, Inc. Complexity 21: 346–360, 2016     

Hopf bifurcation analysis in a multiple delayed innovation diffusion model with Holling II functional response

A nonlinear mathematical model with Holling II functional response describing the dynamics of nonadopter and adopters population in a stage structured innovation diffusion model, which incorporates the evaluation stage (multiple delays), is proposed. Firstly, we study the stability and the existence of periodic solutions via Hopf bifurcation with respect to both delays at the positive equilibrium by analyzing the distribution of the roots of the corresponding exponential characteristic equation obtained through the variational matrix. The direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions are determined with the help of normal form theory and center manifold theorem. Meanwhile, various cases are discussed to examine the effect of different delays on the stability of delayed innovation diffusion system and are also established numerically. It is also observed that the cumulative density of external influences has a significant role in developing maturity stage (adoption stage) in the system. Finally, numerical simulations are carried out to support and supplement the analytical findings.