Effects of population dispersal and impulsive control tactics on pest management

In this paper, we firstly consider a Lotka–Volterra predator–prey model with impulsive constant releasing for natural enemies and a proportion of killing or catching pests at fixed moments, and we have proved that there exists a pest-eradication periodic solution which is globally asymptotically stable. Further, we extend the model for the population to move in a two-patch environment. The effects of population dispersal and impulsive control tactics are investigated, i.e. we chiefly address the question of whether population dispersal is beneficial or detrimental for pest persistence. To do this, some special cases are theoretically investigated and numerical investigations are done for general case. The results indicate that for some ranges of dispersal rates, population dispersal is beneficial to pest control, but for other ranges, it is harmful. These clarify that we can get some new effective pest control strategies by controlling the dispersal rates of pests and natural enemies.     

A study of predator–prey models with the Beddington–DeAnglis functional response and impulsive effect

In this paper, the predator–prey system with the Beddington–DeAngelis functional response is developed, by introducing a proportional periodic impulsive catching or poisoning for the prey populations and a constant periodic releasing for the predator. The Beddington–DeAngelis functional response is similar to the Holling type II functional response but contains an extra term describing mutual interference by predators. This model has the potential to protect predator from extinction, but under some conditions may also lead to extinction of the prey. That is, the system exists a locally stable prey-eradication periodic solution when the impulsive period satisfies an inequality. The condition for permanence is established via the method of comparison involving multiple Liapunov̀ functions. Further, by numerical simulation method the influences of the impulsive perturbations and mutual interference by predators on the inherent oscillation are investigated. With the increasing of releasing for the predator, the system appears a series of complex phenomenon, which include (1) period-doubling, (2) chaos attractor, (3) period-halfing. (4) non-unique dynamics (meaning that several attractors coexist).     

Dynamics of an impulsive control system which prey species share a common predator

In an ecosystem multiple prey species often share a common predator and the interactions between the preys are neutral. In view of these facts and based on a multiple species prey–predator system with Holling IV and II functional responses, an impulsive differential equation to model the process of periodically releasing natural enemies and spraying pesticides at different fixed times for pest control is proposed and investigated. It is proved that there exists a locally asymptotically stable pest-eradication periodic solution under the assumption that the impulsive period is less than some critical value (or the release amount of the predator is greater than another critical value). Permanence conditions are established when the impulsive period is greater than another critical value (or the release amount of the predator is less than some critical value). Numerical results show that the system we consider has more complex dynamics including period solution, quasi-periodic oscillation, chaos, intermittency and crises.     

Attractor information for discrete dynamical systems by means of optimal discrete Galerkin bases

We introduce local adaptive discrete Galerkin bases as a basis set in order to obtain geometrical and topological information about attractors of discrete dynamical systems. The asymptotic behavior of these systems is described by the reconstruction of their attractors in a finite dimensional Euclidean space and by the attractor topological characteristics including the minimal embedding dimension and its local dimension. We evaluate numerically the applicability of our geometrical and topological results by examining two examples: a dissipative discrete system and a nonlinear discrete predator–prey model that includes several types of self-limitation on the prey.     

Extinction and permanence in delayed stage-structure predator–prey system with impulsive effects

Based on the classical stage-structured model and Lotka–Volterra predator–prey model, an impulsive delayed differential equation to model the process of periodically releasing natural enemies at fixed times for pest control is proposed and investigated. We show that the conditions for global attractivity of the ‘pest-extinction’ (‘prey-eradication’) periodic solution and permanence of the population of the model depend on time delay. We also show that constant maturation time delay and impulsive releasing for the predator can bring great effects on the dynamics of system by numerical analysis. As a result, the pest maturation time delay is considered to establish a procedure to maintain the pests at an acceptably low level in the long term. In this paper, the main feature is that we introduce time delay and pulse into the predator–prey (natural enemy-pest) model with age structure, exhibit a new modelling method which is applied to investigate impulsive delay differential equations, and give some reasonable suggestions for pest management.     

一类具有时滞、脉冲效应和阶段结构的捕食者-食饵模型（英文）

A delayed predator-prey model concerning impulsive spraying pesticides and releasing natural enemies is proposed and investigated,in which both the prey and the predator have a history that takes them through two stages:immature and mature.The global attractiveness of the pest-eradication periodic solution is discussed,and sufficient condition is obtained for the permanence of the system.Further,numerical simulations show that there is a characteristic sequence of bifurcations leading to a chaotic dynamics,which implies that the system with constant periodic impulsive perturbations admits rich and complex dynamics.     

The dynamics of pest control pollution model with age structure and time delay

In this paper, by using pollution model and impulsive delay differential equation, we investigate the dynamics of a pest control model with age structure for pest by introducing a constant periodic pesticide input and releasing natural enemies at different fixed moment. We assume only the pests are affected by pesticide. We show that there exists a global attractive pest-extinction periodic solution when the periodic natural enemies release amount μ₁ and pesticide input amount μ₂ are larger than some critical value. Further, the condition for the permanence of the system is also given. By numerical analyses, we also show that constant maturation time delay, pulse pesticide input and pulse releasing of the natural enemies can bring obvious effects on the dynamics of system. We believe that the results will provide reliable tactic basis for the practical pest management.     

An integrated pest management model with delayed responses to pesticide applications and its threshold dynamics

Pulse-like pest management actions such as spraying pesticides and killing a pest instantly and the release of natural enemies at critical times can be modelled with impulsive differential equations. In practice, many pesticides have long-term residual effects and, also, both pest and natural enemy populations may have delayed responses to pesticide applications. In order to evaluate the effects of the duration of the residual effectiveness of pesticides and of delayed responses to pesticides on a pest management strategy, we developed novel mathematical models. These combine piecewise-continuous periodic functions for chemical control with pulse actions for releasing natural enemies in terms of fixed pulse-type actions and unfixed pulse-type actions. For the fixed pulse-type model, the stability threshold conditions for the pest eradication periodic solution and permanence of the model are derived, and the effects of key parameters including killing efficiency rate, decay rate, delayed response rate, number of pesticide applications and number of natural enemy releases on the threshold values are discussed in detail. The results indicate that there exists an optimal releasing period or an optimal number of pesticide applications which maximizes the threshold value. For unfixed pulse-type models, the effects of the killing efficiency rate, decay rate and delayed response rate on the pest outbreak period, and the frequency of control actions are also investigated numerically.     

Modeling of chaotic motion of gyrostats in resistant environment on the base of dynamical systems with strange attractors

A chaotic motion of gyrostats in resistant environment is considered with the help of well known dynamical systems with strange attractors: Lorenz, Rössler, Newton–Leipnik and Sprott systems. Links between mathematical models of gyrostats and dynamical systems with strange attractors are established. Power spectrum of fast Fourier transformation, gyrostat longitudinal axis vector hodograph and Lyapunov exponents are find. These numerical techniques show chaotic behavior of motion corresponding to strange attractor in angular velocities phase space. Cases for perturbed gyrostat motion with variable periodical inertia moments and with periodical internal rotor relative angular moment are considered; for some cases Poincaré sections areobtained.     

Analysis of a predator–prey model with Holling II functional response concerning impulsive control strategy

According to biological and chemical control strategy for pest control, we investigate the dynamic behavior of a Holling II functional response predator–prey system concerning impulsive control strategy-periodic releasing natural enemies and spraying pesticide at different fixed times. By using Floquet theorem and small amplitude perturbation method, we prove that there exists a stable pest-eradication periodic solution when the impulsive period is less than some critical value. Further, the condition for the permanence of the system is also given. Numerical results show that the system we consider can take on various kinds of periodic fluctuations and several types of attractor coexistence and is dominated by periodic, quasiperiodic and chaotic solutions, which implies that the presence of pulses makes the dynamic behavior more complex. Finally, we conclude that our impulsive control strategy is more effective than the classical one if we take chemical control efficiently.     

Seasonally perturbed prey–predator system with predator-dependent functional response

The effect of seasonality on the prey–predator model with predator-dependent trophic function is investigated analytically as well as numerically. The effect of periodic variations is considered on two different parameters of the system: the growth rate of prey and the death rate of the predators. The two parameters may not be in the same phase. The behavior of the system is simulated and bifurcation diagrams are obtained for different parameters. The results show that seasonality in two different parameters with or without phase difference can give rise to multiple attractors, including chaos, with variations in critical parameters.     

Dynamic analysis of a fractional order prey–predator interaction with harvesting

In recent years, prey–predator models appearing in various fields of mathematical biology have been proposed and studied extensively due to their universal existence and importance. In this paper, we introduce a fractional-order prey–predator model and deals with the mathematical behaviors of the model. The dynamical behavior of the system is investigated from the point of view of local stability. We also carry out a detailed analysis on the stability of equilibrium. Numerical simulations are presented to illustrate the results.     

Stationary distribution and extinction for a food chain chemostat model with random perturbation

In this paper, we study the dynamical behavior of a stochastic food chain chemostat model, in which the white noise is proportional to the variables. Firstly, we prove the existence and uniqueness of the global positive solution. Then by constructing suitable Lyapunov functions, we show the system has a unique ergodic stationary distribution. Furthermore, the extinction of microorganisms is discussed in two cases. In one case, both the prey and the predator species are extinct, and in the other case, the prey species is surviving and the predator species is extinct. Finally, numerical experiments are performed for supporting the theoretical results.     

Models for determining how many natural enemies to release inoculatively in combinations of biological and chemical control with pesticide resistance

Combining biological and chemical control has been an efficient strategy to combat the evolution of pesticide resistance. Continuous releases of natural enemies could reduce the impact of a pesticide on them and the number to be released should be adapted to the development of pesticide resistance. To provide some insights towards this adaptation strategy, we developed a novel pest–natural enemy model considering both resistance development and inoculative releases of natural enemies. Three releasing functions which ensure the extinction of the pest population are proposed and their corresponding threshold conditions obtained. Aiming to eradicate the pest population, an analytic formula for the number of natural enemies to be released was obtained for each of the three different releasing functions, with emphasis on their biological implications. The results can assist in the design of appropriate control strategies and decision-making in pest management.     

Stochastic persistence and stability analysis of a modified Holling–Tanner model

The article aims to study the basic dynamical features of a modified Holling–Tanner prey–predator model with ratio‐dependent functional response. We have proved the global existence of the solution for the deterministic model. The parametric restriction for persistence of both species is also obtained along with the proof of local asymptotic stability of the interior equilibrium point(s). Conditions for local bifurcations of interior equilibrium points are provided. The global dynamic behavior is examined thoroughly with supportive numerical simulation results. Next, we have formulated the stochastic model by perturbing the intrinsic growth rates of prey and predator populations with white noise terms. The existence uniqueness of solutions for stochastic model is established. Further, we have derived the parametric restrictions required for the persistence of the stochastic model. Finally, we have discussed the stochastic stability results in terms of the first and second order moments. Numerical simulation results are provided to support the analytical findings. Copyright © 2012 John Wiley & Sons, Ltd.     

Chaos in periodically forced Holling type IV predator–prey system with impulsive perturbations

The effect of periodic forcing and impulsive perturbations on predator–prey model with Holling type IV functional response is investigated. The periodic forcing is affected by assuming a periodic variation in the intrinsic growth rate of the prey. The impulsive perturbations are affected by introducing periodic constant impulsive immigration of predator. The dynamical behavior of the system is simulated and bifurcation diagrams are obtained for different parameters. The results show that periodic forcing and impulsive perturbation can easily give rise to complex dynamics, including (1) quasi-periodic oscillating, (2) period doubling cascade, (3) chaos, (4) period halfing cascade.     

Hybrid approach for pest control with impulsive releasing of natural enemies and chemical pesticides: A plant–pest–natural enemy model

The agricultural pests can be controlled effectively by simultaneous use (i.e., hybrid approach) of biological and chemical control methods. Also, many insect natural enemies have two major life stages, immature and mature. According to this biological background, in this paper, we propose a three tropic level plant–pest–natural enemy food chain model with stage structure in natural enemy. Moreover, impulsive releasing of natural enemies and harvesting of pests are also considered. We obtain that the system has two types of periodic solutions: plant–pest-extinction and pest-extinction using stroboscopic maps. The local stability for both periodic solutions is studied using the Floquet theory of the impulsive equation and small amplitude perturbation techniques. The sufficient conditions for the global attractivity of a pest-extinction periodic solution are determined by the comparison technique of impulsive differential equations. We analyze that the global attractivity of a pest-extinction periodic solution and permanence of the system are evidenced by a threshold limit of an impulsive period depending on pulse releasing and harvesting amounts. Finally, numerical simulations are given in support of validation of the theoretical findings.     

Comment on “Existence and stability of periodic solution of a Lotka-Volterra predator-prey model with state dependent impulsive effects” [J. Comput. Appl. Math. 224 (2009) 544-555]

In this paper, according to integrated pest management principles, a class of Lotka-Volterra predator-prey model with state dependent impulsive effects is presented. In this model, the control strategies by releasing natural enemies and spraying pesticide at different thresholds are considered. The sufficient conditions for the existence and stability of the positive order-1 periodic solution are given by the Poincaré map and the properties of the LambertW function.     

Approximate Forward Attractors of Non-Autonomous Dynamical Systems

In this paper the forward asymptotical behavior of non-autonomous dynamical systems and their attractors are investigated. Under general conditions, the authors show that every neighborhood of pullback attractor has forward attracting property.     

Dynamics of an intraguild predation food web model with strong Allee effect in the basal prey

Since intraguild predation (IGP) is a ubiquitous and important community module in nature and Allee effect has strong impact on population dynamics, in this paper we propose a three-species IGP food web model consisted of the IG predator, IG prey and basal prey, in which the basal prey follows a logistic growth with strong Allee effect. We investigate the local and global dynamics of the model with emphasis on the impact of strong Allee effect. First, positivity and boundedness of solutions are studied. Then existence and stability of the boundary and interior equilibria are presented and the Hopf bifurcation curve at an interior equilibrium is given. The existence of a Hopf bifurcation curve indicates that if competition between the IG prey and IG predator for the basal resource lies below the curve then the interior equilibrium remains stable, while if it lies above the curve then the interior equilibrium loses its stability. In order to explore the impact of Allee effect, the parameter space is classified into sixteen different regions and, in each region, the number of interior equilibria is determined and the corresponding bifurcation diagrams on the Allee threshold are given. The extinction parameter regions of at least one species and the necessary coexistence parameter regions of all three species are provided. In addition, we explore possible dynamical patterns, i.e., the existence of multiple attractors. By theoretical analysis and numerical simulations, we show that the model can have one (i.e. extinction of all species), two (i.e. bi-stability) or three (i.e. tri-stability) attractors. It is also found by simulations that when there exists a unique stable interior equilibrium, the model may generate multiple attracting periodic orbits and the coexistence of all three species is enhanced as the competition between the IG prey and IG predator for the basal resource is close to the Hopf bifurcation curve from below. Our results indicate that the intraguild predation food web model exhibits rich and complex dynamic behaviors and strong Allee effect in the basal prey increases the extinction risk of not only the basal prey but also the IG prey or/and IG predator.