Dynamics of an intraguild predation food web model with strong Allee effect in the basal prey

Since intraguild predation (IGP) is a ubiquitous and important community module in nature and Allee effect has strong impact on population dynamics, in this paper we propose a three-species IGP food web model consisted of the IG predator, IG prey and basal prey, in which the basal prey follows a logistic growth with strong Allee effect. We investigate the local and global dynamics of the model with emphasis on the impact of strong Allee effect. First, positivity and boundedness of solutions are studied. Then existence and stability of the boundary and interior equilibria are presented and the Hopf bifurcation curve at an interior equilibrium is given. The existence of a Hopf bifurcation curve indicates that if competition between the IG prey and IG predator for the basal resource lies below the curve then the interior equilibrium remains stable, while if it lies above the curve then the interior equilibrium loses its stability. In order to explore the impact of Allee effect, the parameter space is classified into sixteen different regions and, in each region, the number of interior equilibria is determined and the corresponding bifurcation diagrams on the Allee threshold are given. The extinction parameter regions of at least one species and the necessary coexistence parameter regions of all three species are provided. In addition, we explore possible dynamical patterns, i.e., the existence of multiple attractors. By theoretical analysis and numerical simulations, we show that the model can have one (i.e. extinction of all species), two (i.e. bi-stability) or three (i.e. tri-stability) attractors. It is also found by simulations that when there exists a unique stable interior equilibrium, the model may generate multiple attracting periodic orbits and the coexistence of all three species is enhanced as the competition between the IG prey and IG predator for the basal resource is close to the Hopf bifurcation curve from below. Our results indicate that the intraguild predation food web model exhibits rich and complex dynamic behaviors and strong Allee effect in the basal prey increases the extinction risk of not only the basal prey but also the IG prey or/and IG predator.     

Dynamics in a discrete-time predator-prey system with Allee effect

In this paper, dynamics of the discrete-time predator-prey system with Allee effect are investigated in detail. Conditions of the existence for flip bifurcation and Hopf bifurcation are derived by using the center manifold theorem and bifurcation theory, and then further illustrated by numerical simulations. Chaos in the sense of Marotto is proved by both analytical and numerical methods. Numerical simulations included bifurcation diagrams, Lyapunov exponents, phase portraits, fractal dimensions display new and rich dynamical behavior. More specifically, apart from stable dynamics, this paper presents the finding of chaos in the sense of Marotto together with a host of interesting phenomena connected to it. The analytic results and numerical simulations demostrates that the Allee constant plays a very important role for dynamical behavior. The dynamical behavior can move from complex instable states to stable states as the Allee constant increases (within a limited value). Combining the existing results in the current literature with the new results reported in this paper, a more complete understanding of the discrete-time predator-prey with Allee effect is given.     

A discrete-time model with non-monotonic functional response and strong Allee effect in prey

In this paper, we investigate the impact of strong Allee effect on the stability of a discrete-time predator–prey model with a non-monotonic functional response. The dynamics of discrete-time predator–prey models with strong Allee effect is studied earlier. But, the mathematical investigations of predator–prey dynamics in discrete-time set up with Holling type-IV functional response and strong Allee effect in prey are lacking. The proposed model supports the coexistence of two steady states, and the mathematical features of the model are analyzed based on local stability and bifurcation theory. By considering the Allee parameter as the bifurcation parameter, we provide sufficient conditions for the flip and the Neimark–Sacker bifurcations. We observe that Allee parameter plays a significant role in the dynamics of the system.     

An extended discrete Ricker population model with Allee effects

Based on the classical discrete Ricker population model, we incorporate Allee effects by assuming rectangular hyperbola, or Holling-II type functional form, for the birth or growth function and formulate an extended Ricker model. We explore the dynamics features of the extended Ricker model. We obtain domains of attraction for the trivial fixed point. We determine conditions for the existence and stability of positive fixed points and find regions where there exist no positive fixed points, two positive fixed points one of which is stable and two positive fixed points both of which are unstable. We demonstrate that the model exhibits period-doubling bifurcations and investigate the existence and stability of the cycles. We also confirm that Allee effects have stabilization effects, by different measures, through numerical simulations.     

Dynamical analysis in a diffusive predator-prey system with a delay and strong Allee effect

In this paper, we consider the dynamics of a diffusive predator-prey system with strong Allee effect and delayed Ivlev-type functional response. At first, we apply the method of upper-lower solutions and the comparison principle in proving the nonnegativity of the solutions. Then by analyzing the distribution of the eigenvalues, we obtain the bistability of the system and the existence of Hopf bifurcation. Furthermore, by using the center manifolds theory and normal form method, we study the properties of the Hopf bifurcations. Finally, some numerical simulations are carried out for illustrating the theoretical results.     

Dynamics of a Predator-Prey Model with Allee Effect and Herd Behavior

This paper deals with dynamics of a predator-prey model with Allee effect and herd behavior. We first study the stability of non-negative constant solutions for such system. We also establish the existence of Hopf bifurcation solutions for such predator-prey model. The stability and bifurcation direction of Hopf bifurcation solution in the case of spatial homogeneity are further discussed. At the same time, several examples are given by MATLAB. Finally, the numerical simulations of the system are carried out through MATLAB, which intuitively verifies and supplements the theoretical analysis results.     

Transitions in a logistic population model incorporating an Allee effect

In some species, the population may decline to zero; that is, the species becomes extinct if the population falls below a given threshold. This phenomenon is well known as an Allee effect. In most Allee models, the model parameters are constants, and the population tends either to a nonzero limiting state (survival) or to zero (extinction). However, when environmental changes occur, these parameters may be slowly varying functions of time. Then, application of multitiming techniques allows us to construct approximations to the evolving population in cases where the population survives to a slowly varying surviving state and those where the population declines to zero. Here, we investigate the solution of a logistic population model exhibiting an Allee effect, when the carrying capacity and the limiting density interchange roles, via a transition point. We combine multiscaling analysis with local asymptotic analysis at the transition point to obtain an overall expression for the evolution of the population. We show that this shows excellent agreement with the results of numerical computations. Copyright © 2015 John Wiley & Sons, Ltd.     

Dynamics of a delayed predator-prey model with Allee effect and Holling type II functional response

In this paper, a delayed with Holling type II functional response (Beddington-DeAngelis) and Allee effect predator-prey model is considered. The growth of the prey is affected by the parameter $M$, which defines the Allee effect. In addition, the delay $\tau$ also influences the logistic growth of the prey, which can be interpreted as the maturity time or the gestation period. In the study of the characteristic equation, we observe that the delay $\tau$ also depends on the parameter $M$, which affects the dynamics in the prey population. Considering the delay as a bifurcation parameter, the local asymptotic stability of the positive equilibrium is investigated. On the other hand, we find that the system can also suffer a Hopf bifurcation in the positive equilibrium when the delay passes through a sequence of critical values. In particular, we study the direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions, an explicit algorithm is provided applying the normal form theory and center manifold reduction for the functional differential equations. Finally, numerical simulations that support the theoretical analysis are included.     

具有HollingⅡ型功能反应和Allee效应的捕食系统模型

利用计算机模拟方法研究一类离散种群相互作用模型的动态复杂性.通过理论推导建立食饵具有Allee效应和HollingⅡ型功能反应的自治捕食系统模型,用Matlab软件模拟离散种群的生长状态,探索研究参数的变化对种群大小的影响,阐释Allee效应及HollingⅡ型功能反应在种群间相互作用模型中的重要性.研究结果表明:1)当处理时间处于有效区间内时,处理时间越大种群的稳定共存参数域越大;2)Allee效应的引入使种群的动态行为更为复杂,从而增加了捕食者种群的灭绝风险;3)系统受强Allee效应的影响,种群会出现提前分叉现象,如果继续增加Allee效应就会导致种群灭绝;4)强Allee效应更容易使种群趋向灭绝.所得结论在丰富生态学理论的同时,提出了保护生态学的重要依据.     

Multiscaling analysis of a slowly varying single species population model displaying an Allee effect

We consider the growth of a single species population modelled by a logistic equation modified to accommodate an Allee effect, in which the model parameters are slowly varying functions of time. We apply a multitiming technique to construct general approximate expressions for the evolving population in the case where the population survives to a (slowly varying) finite positive limiting state, and that where the population declines to extinction. We show that these expressions give excellent agreement with the results of numerical calculations for particular instances of the changing model parameters. Copyright © 2013 John Wiley & Sons, Ltd.     

Non-autonomous periodic systems with Allee effects

A new class of maps called unimodal Allee maps are introduced. Such maps arise in the study of population dynamics in which the population goes extinct if its size falls below a threshold value. A unimodal Allee map is thus a unimodal map with three fixed points, a zero fixed point, a small positive fixed point, called threshold point, and a bigger positive fixed point, called the carrying capacity. In this paper, the properties and stability of the three fixed points are studied in the setting of non-autonomous periodic dynamical systems or difference equations. Finally, we investigate the bifurcation of periodic systems/difference equations when the system consists of two unimodal Allee maps.     

Optimal harvesting in a predator-prey model with Allee effect and sigmoid functional response

This work deals with the determination of the optimal harvest policy in an open access fishery in which both prey and predator species are subjected to non-selective harvesting.The model is described by autonomous ordinary differential equation systems, the functional response of the predators is Holling type III and the prey growth is affected by the Allee effect. The catch-rate functions are based on the catch per unit effort (CPUE) or Schaefer’s hypothesis.The problem of determining the optimal harvest policy is solved by using Pontryagin’s maximal principle. The problem here studied is to maximize a cost function representing the present value of a continuous time-stream of revenue of the fishery.     

Limit cycles in a Gause‐type predator–prey model with sigmoid functional response and weak Allee effect on prey

The goal of this work is to examine the global behavior of a Gause‐type predator–prey model in which two aspects have been taken into account: (i) the functional response is Holling type III; and (ii) the prey growth is affected by a weak Allee effect. Here, it is proved that the origin of the system is a saddle point and the existence of two limit cycles surround a stable positive equilibrium point: the innermost unstable and the outermost stable, just like with the strong Allee effect. Then, for determined parameter constraints, the trajectories can have different ω ? limit sets. The coexistence of a stable limit cycle and a stable positive equilibrium point is an important fact for ecologists to be aware of the kind of bistability shown here. So, these models are undoubtedly rather sensitive to disturbances and require careful management in applied contexts of conservation and fisheries. Copyright © 2012 John Wiley & Sons, Ltd.     

Pattern formation in a variable diffusion predator–prey model with additive Allee effect

This paper deals with a variable diffusion predator–prey model with additive Allee effect. A good understanding of the existence of steady states is gained for the case  $\sigma =0$. The result shows that the reduce problem has multiple solutions. Moreover, by applying the singular perturbation method, we give a proof of existence of large amplitude solutions when  $\sigma$ is sufficiently small.     

Population models with diffusion, strong Allee effect and constant yield harvesting

We study the steady state distribution of reaction diffusion equations with strong Allee effect type growth and constant yield harvesting (semipositone) in heterogeneous bounded habitats. Assuming the exterior of the habitat is completely hostile, we establish existence results for positive solutions. We also establish a multiplicity result for the non-harvested case. We obtain our results via the method of sub-super solutions.     

Global qualitative analysis of a discrete host‐parasitoid model with refuge and strong Allee effects

In this paper, we discuss the qualitative behavior of a discrete host‐parasitoid model with the host subject to refuge and strong Allee effects. More precisely, we study the local and global asymptotic stability, stable manifolds and unstable manifolds of boundary equilibrium points, existence and unique positive equilibrium point, local and global behavior of the positive equilibrium point, and the uniform persistence for the model with the host subject to the refuge or both refuge and strong Allee effects. It is also proved that the model undergoes a transcritical bifurcation in a small neighborhood of the boundary equilibrium point. Some numerical simulations are given to support our theoretical results. We can obtain that the addition of the refuge may make the parasitoids go extinct while the hosts survive or may stabilize the host‐parasitoid interaction; the addition of both refuge and strong Allee effects has either a negative or positive impact on the coexistence of both populations.     

Allee effects in a discrete-time SIS epidemic model with infected newborns

We extend the framework of Rios-Soto et al. (Contemporary Mathematics, 2006, 410, 297) to include both compensatory (contest competition) and overcompensatory (scramble competition) population dynamics with and without the Allee effect. We compute the basic reproductive number ?₀, and use it to predict the (uniform) persistence or extinction of the infective population, where the population dynamics are compensatory and the Allee effect is either present or absent. We also explore the relationship between the demographic equation and the epidemic process, where the total population dynamics are overcompensatory. In particular, we show that the demographic dynamics drive both the susceptible and infective dynamics. This is in contrast to the recent observations of Franke and Yakubu, that the demographic dynamics can be chaotic while the infective dynamics are oscillatory and non-chaotic in periodically-forced SIS epidemic models (Mathematical Biosciences, 2006, 204, 68).