Routes to bursting in a periodically driven oscillator

The dynamical behaviors of a periodic excited oscillator with multiple time scales in the form that order gap exists between the frequency of the excitation and the natural frequency, are investigated in this Letter. By regarding the whole excitation term as a parameter, bifurcation sets are derived, which divide the generalized parameter space into several regions corresponding to different kinds of dynamics. Different types of bursting phenomena, such as fold/Hopf bursting, fold/Hopf/homoclinic bursting and Hopf/homoclinic bursting, are presented, the mechanism of which is obtained based on the bifurcations of the generalized autonomous system as well as the introduction of the so-called transformed phase portraits. Furthermore, the evolution of the bursting is discussed in details, in which one may find that when the two limit cycles caused by the Hopf bifurcations of the two related equilibrium points interact with each other, homoclinic bifurcation may occur, leading to the merge of the two cycles to form a large amplitude cycle. The homoclinic bifurcation may cause the two asymmetric bursters to merge into a symmetric enlarged burster, in which the large amplitude of the spiking state agrees well with the amplitude of the cycle caused by the homoclinic bifurcation.     

两时间尺度下非光滑广义蔡氏电路系统的簇发振荡机理

通过引入周期变化的电流源并选择适当参数, 使得周期激励频率与系统固有频率之间存在量级差距, 建立了两时间尺度即快慢耦合非光滑广义蔡氏电路模型. 基于相应的广义自治系统, 考察了其不同区域中的平衡态及其稳定性, 得到了不同分岔行为及其相应的临界条件. 同时, 利用广义Clarke导数得到的广义Jacobian矩阵, 探讨了系统轨迹穿越非光滑分界面时的各种非常规分岔模式, 进而结合广义相图, 深入分析了Fold/Fold周期簇发振荡以及Fold/Hopf周期簇 发振荡两种典型的周期簇发行为及其相应的分岔机制. 关键词： 非光滑 广义蔡氏电路 两时间尺度 分岔机制     

Bursting phenomena as well as the bifurcation mechanism in controlled Lorenz oscillator with two time scales

A controlled Lorenz model with fast-slow effect has been established, in which there exist order gap between the variables associated with the controller and the original Lorenz oscillator, respectively. The conditions of fold bifurcation as well as Hopf bifurcation for the fast subsystem are derived to investigate the mechanism of the behaviors of the whole system. Two cases in which the equilibrium points of the fast subsystem behave in different characteristics have been considered, leading to different dynamical evolutions with the change of coupling strength. Several types of bursting phenomena, such as fold/fold burster, fold/Hopf burster, near-fold/Hopf burster, fold/near-Hopf buster have been observed. Theoretical analysis shows that the bifurcations points which connect the quiescent state and the repetitive spiking state agree well with the turning points of the trajectories of the bursters. Furthermore, the mechanism of the period-adding bifurcations, resulting in the rapid change of the period of the movements, is presented.     

Bursting phenomena as well as the bifurcation mechanism in a coupled BVP oscillator with periodic excitation

We explore the complicated bursting oscillations as well as the mechanism in a high-dimensional dynamical system.By introducing a periodically changed electrical power source in a coupled BVP oscillator, a fifth-order vector field with two scales in frequency domain is established when an order gap exists between the natural frequency and the exciting frequency.Upon the analysis of the generalized autonomous system, bifurcation sets are derived, which divide the parameter space into several regions associated with different types of dynamical behaviors. Two typical cases are focused on as examples,in which different types of bursting oscillations such as sub Hopf/sub Hopf burster, sub Hopf/fold-cycle burster, and doublefold/fold burster can be observed. By employing the transformed phase portraits, the bifurcation mechanism of the bursting oscillations is presented, which reveals that different bifurcations occurring at the transition between the quiescent states(QSs) and the repetitive spiking states(SPs) may result in different forms of bursting oscillations. Furthermore, because of the inertia of the movement, delay may exist between the locations of the bifurcation points on the trajectory and the bifurcation points obtained theoretically.     

慢变控制下Chen系统的复杂行为及其机理

由于Chen系统的控制分析大都是基于同一时间尺度,而两时间尺度耦合问题的相关研究基本上局限于单维慢变量情形.本文探讨了基于慢时间尺度上的Duffing振子,即含有两维慢子系统控制下Chen系统的动力学演化过程.给出了诸如对称式fold/fold、对称式fold/Hopf、对称式homoclinic/homoclinic等不同形式的簇发振荡行为,并揭示了其相应的产生机制,指出慢子系统中两维慢变量的相互影响导致系统产生了类似于周期激励下的簇发行为.     

多平衡态下簇发振荡产生机理及吸引子结构分析

以周期激励下受控Lorenz模型为例, 考察了多平衡态共存下激励频率与系统固有频率之间存在量级差距也即存在频域上的不同尺度时的耦合效应. 由于激励频率远小于系统的固有频率, 因此将整个激励项视为慢变参数, 分析随慢变参数变化下的各种分岔模式及其相应的分岔行为, 指出在一定条件下, 不同平衡点会产生Hopf分岔和fold分岔. 根据分岔条件的不同, 给出了两种典型情况下的簇发振荡, 并通过引入转换相图, 揭示了不同簇发的产生机理, 指出多平衡态和多种分岔共存不仅会导致沉寂态和激发态的多样性, 而且会使得不同沉寂态和激发态之间存在着不同的转换形式.     

铂族金属氧化过程中的簇发振荡及其诱发机理

铂族金属表面氧化过程是典型的多相催化反应之一, 具有广泛的应用背景及丰富的振荡行为, 因此深入研究铂族金属的氧化中的物理及化学过程具有重要的理论意义及工程应用前景. 通过对铂族金属CO的氧化过程中实测数据的回归分析, 建立了不同尺度耦合解析动力学理论模型. 通过对平衡态的稳定性分析, 指出在一定条件下稳态解会由鞍-结同宿轨道分岔导致周期振荡. 当快子系统产生Hopf分岔时, 该周期振荡会进一步演化为两尺度耦合的周期簇发振荡, 即N^k振荡, 并由加周期分岔使得系统处于激发态的时间显著增加.在此基础上, 利用分岔理论进一步分析了周期簇发及加周期分岔的产生机理, 揭示了周期簇发中沉寂态和激发态相互转化时的不同分岔模式.     

Bursting oscillations in a hydro-turbine governing system with two time scales

A fast-slow coupled model of the hydro-turbine governing system(HTGS)is established by introducing frequency disturbance in this paper.Based on the proposed model,the performances of two time scales for bursting oscillations in the HTGS are investigated and the effect of periodic excitation of frequency disturbance is analyzed by using the bifurcation diagrams,time waveforms and phase portraits.We find that stability and operational characteristics of the HTGS change with the value of system parameter k_d.Furthermore,the comparative analyses for the effect of the bursting oscillations on the system with different amplitudes of the periodic excitation a are carried out.Meanwhile,we obtain that the relative deviation of the mechanical torque mt rises with the increase of a.These methods and results of the study,combined with the performance of two time scales and the fast-slow coupled engineering model,provide some theoretical bases for investigating interesting physical phenomena of the engineering system.     

Bursting Ca2+ Oscillations and Synchronization in Coupled Cells

A mathematical model proposed by Grubelnk et al. [Biophys. Chem. 94 （2001） 59] is employed to study the physiological role of mitochondria and the cytosolic proteins in generating complex Ca^2＋ oscillations, lntracellulax bursting calcium oscillations of point-point, point-cycle and two-folded limit cycle types are observed and explanations are given based on the fast/slow dynamical analysis, especially for point-cycle and two-folded limit cycle types, which have not been reported before. Furthermore, synchronization of coupled bursters of Ca^2＋ oscillations via gap junctions and the effect of bursting types on synchronization of coupled cells are studied. It is argued that bursting oscillations of point-point type may be superior to achieve synchronization than that of point cycle type.     

Symmetric bursting of focus-focus type in the controlled Lorenz system with two time scales

By employing a special feedback controlling scheme, a hyperchaotic Lorenz system with the structure of two time scales is constructed. Two kinds of bursting phenomena, symmetric fold/fold bursting and symmetric sub-Hopf/sub-Hopf bursting, can be observed in this system. Their respective dynamical behaviors are investigated by means of slow-fast analysis. In particular, symmetric fold/fold bursting is of focus-focus type, namely, both the up-state and the down-state are stable focus, which is different from the usual fold/fold bursting; Symmetric sub-Hopf/sub-Hopf bursting is also of focus-focus type, which has not been reported in previous work. Furthermore, phase plane analysis has been introduced to explore the evolution details of the fast subsystem for symmetric sub-Hopf/sub-Hopf bursting. With the variation of the parameter, symmetric sub-Hopf/sub-Hopf bursting can evolve to symmetric chaotic bursting or even hyperchaos.     

快慢型超混沌Lorenz系统分析

讨论了快慢两时间尺度下超混沌Lorenz系统原点的稳定性问题,分析了原点的Hopf分岔,包括Hopf分岔的存在性,分岔方向以及分岔周期解的稳定性等问题,并用数值例子对所得到的结果加以验证.在一定的参数条件下,快慢系统会产生对称簇发并能达到超混沌状态.基于快慢分析法,揭示了对称簇发中沉寂态与激发态相互转迁的不同分岔模式,并进一步分析了耦合强度对慢过效应的影响. 关键词： 超混沌Lorenz系统 Hopf分岔 对称式fold/subHopf簇发 慢过效应     

可兴奋性细胞混沌放电区间的识别机理

在神经起步点记录到加周期分岔过程的生理实验数据，在对此分岔过程中位于周期n爆发 和周期(n+1)爆发之间的混沌的峰峰间期数据检测不稳定的周期轨道时，发现从靠近周期 n爆发的混沌的峰峰间期数据中，可以检测出不稳定的周期n轨道；而从靠近周期(n+1)爆 发的混沌的峰峰间期数据中，不仅可以检测出不稳定的周期(n+1)轨道，还可以检测出不稳 定的周期n轨道.针对该现象，借助于Sherman建议的胰腺β细胞模型，从非线性动力 学角度给出了理论解释.指明了由鞍结分岔和倍周期分岔分别产生第一类阵发和第三类阵发 为出现该 关键词： 峰峰间期 不稳定的周期轨道 鞍结分岔 倍周期分岔     

具有多分界面的非线性电路中的非光滑分岔

本文分析了具有多分界面的非线性电路在不同时间尺度下的快慢动力学行为. 在一定的参数条件下,系统的周期解为簇发解,表现出明显的快慢效应. 根据状态变量变化的快慢,把全系统划分为快子系统和慢子系统两组. 根据快慢分析法将慢变量看作快子系统的控制参数,分析了快子系统的平衡点在向量场不同区域内的稳定性. 非光滑系统的分岔与向量场的分界面密切相关,对于具有快慢效应的两时间尺度非光滑系统,快子系统的分岔则取决于分界面两侧平衡点的性质. 通过在临界面引入广义Jacobi矩阵,讨论了快子系统非光滑分岔的类型,即多次穿越分 关键词： 非线性电路 多分界面 非光滑分岔 快慢效应     

三个不同时间尺度的电耦合模型的组合簇放电

胰岛中间隙连接的胰腺β细胞的簇放电行为对胰岛素分泌起着重要的作用. 本文利用了最小的phantom 簇放电模型, 研究两个电耦合胰腺β细胞具有簇同步的组合簇放电, 其膜电位表现出一个长簇和几个短簇组成的放电簇集和振幅先减小后增大的小振幅阈下振荡的相互转迁. 在两个慢变量和快的膜电位的三维空间中, 分别考虑了中慢变量和慢慢变量作为分岔参数的多层次的快慢动力学分析, 研究这两个时间尺度不同的慢变量如何共同或单独地控制着这种组合簇放电的复杂动力学行为. 特别地, 探讨了耦合强度引起的组合簇放电的每个簇集中短簇个数变化的内在机理. 关键词： 电耦合 具有不同时间尺度的慢变量 组合簇放电 快慢动力学分析     

神经元电活动不同节律模式的几种变化过程

利用神经元Chay模型,对实验中观察到的三种放电节律模式序列进行数值模拟,并应用余维1极限环分岔分析研究了其产生机理.首先考虑的是周期性放电模式的变化过程;其次,具有不同表象的一种超临界和一种亚临界倍周期簇放电序列产生并导致混沌现象的出现,然后以不同的方式转迁到逆超临界倍周期峰放电序列;最后研究无混沌的加周期簇放电序列,得出加周期分岔仅是一种与倍周期分岔密切相关的分岔现象.     

计算相位响应曲线的方波扰动直接算法

通过相位响应曲线可对具有极限环周期运动的动力系统的性质有更为深入的理解.神经元是一个典型的动力系统,因此相位响应曲线提供了一种研究神经元重复周期放电行为的新思路.本文提出一种求解相位响应曲线的方法,即方波扰动的直接算法,通过Hodgkin-Huxley,Fitz Hugh-Nagumo,Morris-Lecar和Hindmarsh-Rose神经元模型验证该算法可计算周期峰放电、周期簇放电的相位响应曲线.该算法克服了其他算法在运用过程中的局限性.利用该算法计算结果表明:周期峰放电的相位响应曲线类型是由其分岔类型所决定;在Morris-Lecar模型中发现一种开始于Hopf分岔终止于鞍点同宿轨道分岔的阈上周期振荡,其相位响应曲线属于第二类型.通过大量的相位响应曲线的计算发现相位响应的相对大小及正负性仅取决于扰动所施加的时间,而且周期簇放电的相位响应曲线比周期峰放电的相位响应曲线更为复杂.     

高维广义蔡氏电路中的快慢动力学行为及其分岔分析

讨论了四阶广义蔡氏电路在两时间尺度下的动力学行为. 由数值模拟得到了系统在不同参数条件下的周期簇发解和混沌吸引子. 通过引入快慢分析法,从分岔的角度,以周期解为例, 对系统动力学行为产生的机理及其演化规律进行了理论分析和解释, 其结论与数值计算的结果基本符合.     

Bifurcation Analysis and Transition Mechanism in a Modified Model of Ca~(2+) Oscillations

Some new elements are introduced into a mathematical model of intracellular calcium oscillations, which make it particularly suitable for the study of bifurcation. In addition to generating regular oscillations, such a modified model can be used to reproduce the burst discharges similar to those recorded in experiments and to describe two new types of oscillatory phenomena. By means of a fast/slow dynamical analysis, we explore the bifurcation and transition mechanisms associated with two types of bursters due to changes in the interaction of two slow variables with different timescales.     

Pre-B?tzinger复合体的从簇到峰放电的同步转迁及分岔机制

Pre-Bötzinger复合体是兴奋性耦合的神经元网络,通过产生复杂的放电节律和节律模式的同步转迁参与调控呼吸节律.本文选用复杂簇和峰放电节律的单神经元数学模型构建复合体模型,仿真了与生物学实验相关的多类同步节律模式及其复杂转迁历程,并利用快慢变量分离揭示了相应的分岔机制.当初值相同时,随着兴奋性耦合强度的增加,复合体模型依次表现出完全同步的“fold/homoclinic”,“subHopf/subHopf”簇放电和周期1峰放电.当初值不同时,随耦合强度增加,表现为由“fold/homoclinic”,到“fold/fold limit cycle”、到“subHopf/subHopf”与“fold/fold limit cycle”的混合簇放电、再到“subHopf/subHopf”簇放电的相位同步转迁,最后到反相同步周期1峰放电.完全(同相)同步和反相同步的周期1节律表现出了不同分岔机制.反相峰同步行为给出了与强兴奋性耦合容易诱发同相同步这一传统观念不同的新示例.研究结果给出了preBötzinger复合体的从簇到峰放电节律的同步转迁规律及复杂分岔机制,反常同步行为丰富了非线性动力学的内涵.     

Bursting dynamics in parametrically driven memristive Jerk system

Compared with general nonlinear systems, multi-time scale system has complex bursting dynamics and has received widespread attention. A memristor-based Jerk system with parametric excitation is proposed in this study. As the selected excitation frequency is far less than the natural frequency, implying the existence of an order gap between the excitation frequency and the natural one, the system can be considered as a classic fast-slow system with two timescales. In our system, when the slow-varying parameters periodically pass through the critical pitchfork bifurcation point periodically, a distinct time delay behavior can be observed. Complex bursting oscillations induced by the delayed pitchfork are revealed with different excitation amplitudes. By virtue of the fast-slow analysis method, the corresponding generation mechanisms are discussed by the transformed phase portraits, the time series, and the phase portraits. As the delay time interval induced by the pitchfork bifurcation is dependant not only on the excitation amplitude, but also on the excitation frequency, some excitation frequency related bursting patterns are also considered in our study. Finally, numerical simulations are provided to verify the validity of the study.