Influence of discrete delay on pattern formation in a ratio-dependent prey–predator model

In this paper we explore how the two mechanisms, Turing instability and Hopf bifurcation, interact to determine the formation of spatial patterns in a ratio-dependent prey–predator model with discrete time delay. We conduct both rigorous analysis and extensive numerical simulations. Results show that four types of patterns, cold spot, labyrinthine, chaotic as well as mixture of spots and labyrinthine can be observed with and without time delay. However, in the absence of time delay, the two aforementioned mechanisms have a significant impact on the emergence of spatial patterns, whereas only Hopf bifurcation threshold is derived by considering the discrete time delay as the bifurcation parameter. Moreover, time delay promotes the emergence of spatial patterns via spatio-temporal Hopf bifurcation compared to the non-delayed counterpart, implying the destabilizing role of time delay. In addition, the destabilizing role is prominent when the magnitude of time delay and the ratio of diffusivity are comparatively large.     

Multiple periodic solutions of a ratio-dependent predator–prey model

A delayed ratio-dependent predator–prey model with non-monotone functional response is investigated in this paper. Some new and interesting sufficient conditions are obtained for the global existence of multiple positive periodic solutions of the ratio-dependent model. Our method is based on Mawhin’s coincidence degree and some estimation techniques for the a priori bounds of unknown solutions to the equation Lx = λNx. An example is represented to illustrate the feasibility of our main result.     

Hopf bifurcations in a predator–prey system with a discrete delay and a distributed delay

A delayed Lotka–Volterra two-species predator–prey system with discrete hunting delay and distributed maturation delay for the predator population described by an integral with a strong delay kernel is considered. By linearizing the system at the positive equilibrium and analyzing the associated characteristic equation, the asymptotic stability of the positive equilibrium is investigated and Hopf bifurcations are demonstrated. It is found that under suitable conditions on the parameters the positive equilibrium is asymptotically stable when the hunting delay is less than a certain critical value and unstable when the hunting delay is greater than this critical value. Meanwhile, according to the Hopf bifurcation theorem for functional differential equations (FDEs), we find that the system can also undergo a Hopf bifurcation of nonconstant periodic solution at the positive equilibrium when the hunting delay crosses through a sequence of critical values. In particular, by applying the normal form theory and the center manifold reduction for FDEs, an explicit algorithm determining the direction of Hopf bifurcations and the stability of bifurcating periodic solutions occurring through Hopf bifurcations is given. Finally, to verify our theoretical predictions, some numerical simulations are also included at the end of this paper.     

Effects of prey refuge on a ratio-dependent predator–prey model with stage-structure of prey population

In this paper, a stage-structured predator–prey model is proposed and analyzed to study how the type of refuges used by prey population influences the dynamic behavior of the model. Two types of refuges: those that protect a fixed number of prey and those that protect a constant proportion of prey are considered. Mathematical analyses with regard to positivity, boundedness, equilibria and their stabilities, and bifurcation are carried out. Persistence condition which brings out the useful relationship between prey refuge parameter and maturation time delay is established. Comparing the conclusions obtained from analyzing properties of two types of refuges using by prey, we observe that value of maturation time at which the prey population and hence predator population go extinct is greater in case of refuges which protect a constant proportion of prey.     

A ratio-dependent predator–prey model with diffusion

This paper deals with the existence and nonexistence of nonconstant positive steady-state solutions to a ratio-dependent predator–prey model with diffusion and with the homogeneous Neumann boundary condition. We demonstrate that there exists $a_0(b)$ satisfying $0<a_0(b)<m_1$ for $0<b<m_1$, such that if $0<b<m_1$ and $a_0(b)<a<m_1$, then the diffusion can create nonconstant positive steady-state solutions; whereas the diffusion cannot do provided $a>m_1$.     

Stability and bifurcation of a stage-structured predator–prey model with both discrete and distributed delays

This paper concerns with a new delayed predator–prey model with stage structure on prey, in which the immature prey and the mature prey are preyed by predator and the delay is the length of the immature stage. Mathematical analysis of the model equations is given with regard to invariance of non-negativity, boundedness of solutions, permanence and global stability and nature of equilibria. Our work shows that the stage structure on the prey is one of the important factors that affect the extinction of the predator, and the predation on immature prey is a cause of periodic oscillation of population and can make the behaviors of the system more complex. The predation on the immature and mature prey brings both positive and negative effects on the permanence of the predator, if ignore the predation on immature prey in the system, the stage-structure on prey brings only negative effect on the permanence of the predator.     

Qualitative behaviour of a ratio-dependent predator–prey system

This paper deals with the qualitative properties of an autonomous system of differential equations, modeling ratio-dependent predator–prey interactions.This model differs from traditional ratio dependent models essentially in the predator mortality term, the death rate of the predator is not constant but instead increases when there is overcrowding.We incorporate delay(s) into the system. The most important observation is that as the delay(s) is (are) increased the originally asymptotic stable interior equilibrium loses its stability. Furthermore at a certain critical value a Hopf bifurcation takes place: small amplitude periodic solutions arise.     

Global dynamic behavior of a predator–prey model under ratio-dependent state impulsive control

This paper studies the global dynamic behavior of a prey–predator model with square root functional response under ratio-dependent state impulsive control strategy. It is shown that the boundary equilibrium point of the controlled system is globally asymptotically stable. An order-k periodic orbit is obtained by employing the Brouwer’s fixed point theorem. Furthermore, the critical values are determined for the existence of orbitally asymptotically stable order-1 and order-2 periodic orbits in finite time. These critical values play an important role in determining different kinds of order-k periodic orbits and can also be used for designing the control parameters to obtain the desirable dynamic behavior of the controlled prey–predator system. Moreover, it is found that the local equilibrium point is also globally asymptotically stable under the control strategy. Numerical examples are provided to validate the effectiveness and feasibility of the theoretical results.     

Pattern formation of a predator–prey model

In this paper, we analyze the spatial pattern of a predator–prey system. We get the critical line of Hopf and Turing bifurcation in a spatial domain. In particular, the exact Turing domain is given. Also we perform a series of numerical simulations. The obtained results reveal that this system has rich dynamics, such as spotted, stripe and labyrinth patterns, which shows that it is useful to use the reaction–diffusion model to reveal the spatial dynamics in the real world.     

Chaotic dynamics of a discrete prey–predator model with Holling type II

A discrete-time prey–predator model with Holling type II is investigated. For this model, the existence and stability of three fixed points are analyzed. The bifurcation diagrams, phase portraits and Lyapunov exponents are obtained for different parameters of the model. The fractal dimension of a strange attractor of the model was also calculated. Numerical simulations show that the discrete model exhibits rich dynamics compared with the continuous model, which means that the present model is a chaotic, and complex one.     

Bifurcation and chaos in a ratio-dependent predator–prey system with time delay

In this paper, a ratio-dependent predator–prey model with time delay is investigated. We first consider the local stability of a positive equilibrium and the existence of Hopf bifurcations. By using the normal form theory and center manifold reduction, we derive explicit formulae which determine the stability, direction and other properties of bifurcating periodic solutions. Finally, we consider the effect of impulses on the dynamics of the above time-delayed population model. Numerical simulations show that the system with constant periodic impulsive perturbations admits rich complex dynamic, such as periodic doubling cascade and chaos.     

Periodicity in a nonlinear discrete predator–prey system with state dependent delays

With the help of a continuation theorem based on Gaines and Mawhin's coincidence degree, easily verifiable criteria are established for the global existence of positive periodic solutions of the following nonlinear discrete state dependent delays predator–prey systemwhere $a_i,c_j,d_i:Z\to R^{+}$ are positive $\omega$-periodic, $\omega$ is a fixed positive integer. $b_1,b_2:Z\to R^{+}$ are $\omega$-periodic and ${\sum }_{k=0}^{\omega -1}{b}_i(k)>0$. ${\tau }_i,{\sigma }_j,{\rho }_i:Z\times R\times R\to R(i=1,2,\dots ,n,j=1,2,\dots ,m)$ are $\omega$-periodic with respect to their first arguments, respectively. ${\alpha }_i,{\beta }_j,{\gamma }_i(i=1,2,\dots ,n,j=1,2,\dots ,m)$ are positive constants.     

Dynamics of a stage-structured predator–prey model with prey impulsively diffusing between two patches

In this work, we propose a stage-structured predator–prey model, with prey impulsively diffusing between two patches. Using the discrete dynamical system determined by the stroboscopic map, we obtain a predator-extinction periodic solution. Further, the predator-extinction periodic solution is globally attractive. By the theory on the delay and impulsive differential equation, we prove that the investigated system is permanent. Our results indicate that the discrete time delay has influence to the dynamical behaviors of the investigated system.     

A remark on a stochastic predator–prey system with time delays

An autonomous stochastic predator–prey model with time delays is investigated. Almost sufficient and necessary conditions for stability in the mean and extinction of each population are established. Numerical simulations are introduced to support the results.     

A discrete prey–predator model preserving the dynamics of a structurally unstable Lotka–Volterra model

Leslie's method to construct a discrete two dimensional dynamical system dynamically consistent with the Lotka–Volterra type of competing two species ordinary differential equations is applied in a newly extended manner for the Lotka–Volterra prey–predator system which is structurally unstable. We show that, independently of the time step size, the derived discrete prey–predator system is dynamically consistent with the continuous counterpart, keeping the nature of neutrally stable periodic orbit. Further, we show that the extended method to construct the discrete prey–predator system can provide a dynamically consistent model also for the logistic Lotka–Volterra one.     

Stability and Hopf bifurcation of a diffusive predator–prey model with predator saturation and competition

This article discusses a predator–prey system with predator saturation and competition functional response. The local stability, existence of a Hopf bifurcation at the coexistence equilibrium and stability of bifurcating periodic solutions are obtained in the absence of diffusion. Further, we discuss the diffusion-driven instability, Hopf bifurcation for corresponding diffusion system with zero flux boundary condition and Turing instability region regarding the parameters are established. Finally, numerical simulations supporting the theoretical analysis are also included.