Spatial resonance and Turing–Hopf bifurcations in the Gierer–Meinhardt model

Gierer–Meinhardt system as a molecularly plausible model has been proposed to formalize the observation for pattern formation. In this paper, the Gierer–Meinhardt model without the saturating term is considered. By the linear stability analysis, we not only give out the conditions ensuring the stability and Turing instability of the positive equilibrium but also find the parameter values where possible Turing–Hopf and spatial resonance bifurcation can occur. Then we develop the general algorithm for the calculations of normal form associated with codimension-2 spatial resonance bifurcation to better understand the dynamics neighboring of the bifurcating point. The spatial resonance bifurcation reveals the interaction of two steady state solutions with different modes. Numerical simulations are employed to illustrate the theoretical results for both the Turing–Hopf bifurcation and spatial resonance bifurcation. Some expected solutions including stable spatially inhomogeneous periodic solutions and coexisting stable spatially steady state solutions evolve from Turing–Hopf bifurcation and spatial resonance bifurcation respectively.     

Stability and bifurcation analysis of a nutrient-phytoplankton model with time delay

We proposed a nutrient-phytoplankton interaction model with a discrete and distributed time delay to provide a better understanding of phytoplankton growth dynamics and nutrient-phytoplankton oscillations induced by delay. Standard linear analysis indicated that delay can induce instability of a positive equilibrium via Hopf bifurcation. We derived the conditions guaranteeing the existence of Hopf bifurcation and tracked its direction and the stability of the bifurcating periodic solutions. We also obtained the sufficient conditions for the global asymptotic stability of the unique positive steady state. Numerical analysis in the fully nonlinear regime showed that the stability of the positive equilibrium is sensitive to changes in delay values under select conditions. Numerical results were consistent with results predicted by linear analysis.     

Bifurcation analysis of a predator–prey model with memory-based diffusion

In this paper, we investigate the predator–prey model equipped with Fickian diffusion and memory-based diffusion of predators. The stability and bifurcation analysis explores the impacts of the memory-based diffusion and the averaged memory period on the dynamics near the positive steady state. Specifically, when the memory-based diffusion coefficient is less than a critical value, we show that the stability of the positive steady state can be destabilized as the average memory period increases, which leads to the occurrence of Hopf bifurcations. Moreover, we also analyze the bifurcation properties using the central manifold theorem and normal form theory. This allows us to prove the existence of stable spatially inhomogeneous periodic solutions arising from Hopf bifurcation. In addition, the sufficient and necessary conditions for the occurrence of stability switches are also provided.     

Hopf bifurcation in a delayed reaction–diffusion–advection population model

In this paper, we investigate a reaction–diffusion–advection model with time delay effect. The stability/instability of the spatially nonhomogeneous positive steady state and the associated Hopf bifurcation are investigated when the given parameter of the model is near the principle eigenvalue of an elliptic operator. Our results imply that time delay can make the spatially nonhomogeneous positive steady state unstable for a reaction–diffusion–advection model, and the model can exhibit oscillatory pattern through Hopf bifurcation. The effect of advection on Hopf bifurcation values is also considered, and our results suggest that Hopf bifurcation is more likely to occur when the advection rate increases.     

Bifurcation analysis in a diffusive Logistic population model with two delayed density-dependent feedback terms

The present paper is concerned with a diffusive population model of Logistic type with an instantaneous density-dependent term and two delayed density-dependent terms and subject to the zero-Dirichlet boundary condition. By regarding the delay as the bifurcation parameter and analyzing in detail the associated eigenvalue problem, the local asymptotic stability and the existence of Hopf bifurcation for the sufficiently small positive steady state solution are shown. It is found that under the suitable condition, the positive steady state solution of the model will become ultimately unstable after a single stability switch (or change) at a certain critical value of delay through a Hopf bifurcation. However, under the other condition, the positive steady state solution of the model will become ultimately unstable after multiple stability switches at some certain critical values of delay through Hopf bifurcations. In addition, the direction of the above Hopf bifurcations and the stability of the bifurcating periodic solutions are analyzed by means of the center manifold theory and normal form method for partial functional differential equations. Finally, in order to illustrate the correction of the obtained theoretical results, some numerical simulations are also carried out.     

Existence of periodic travelling waves solutions in predator prey model with diffusion

This paper deals with the qualitative analysis of the travelling waves solutions of a reaction diffusion model that refers to the competition between the predator and prey with modified Leslie–Gower and Holling type II schemes. The well posedeness of the problem is proved. We establish sufficient conditions for the asymptotic stability of the unique nontrivial positive steady state of the model by analyzing roots of the forth degree exponential polynomial characteristic equation. We also prove the existence of a Hopf bifurcation which leads to periodic oscillating travelling waves by considering the diffusion coefficient as a parameter of bifurcation. Numerical simulations are given to illustrate the analytical study.     

Hopf bifurcation of an age-structured compartmental pest-pathogen model

This paper investigates an age-structured compartmental pest-pathogen model by using the theory of integrated semigroup. We study the stability of the steady state of the model by analyzing the associated characteristic transcendental equation. It is shown that Hopf bifurcation occurs at a positive steady state as bifurcating parameter passes a sequence of critical values.     

Bifurcation analysis in a delayed diffusive Nicholson’s blowflies equation

The dynamics of a diffusive Nicholson’s blowflies equation with a finite delay and Dirichlet boundary condition have been investigated in this paper. The occurrence of steady state bifurcation with the changes of parameter is proved by applying phase plane ideas. The existence of Hopf bifurcation at the positive equilibrium with the changes of specify parameters is obtained, and the phenomenon that the unstable positive equilibrium state without dispersion may become stable with dispersion under certain conditions is found by analyzing the distribution of the eigenvalues. By the theory of normal form and center manifold, an explicit algorithm for determining the direction of the Hopf bifurcation and stability of the bifurcating periodic solutions are derived.     

Multiple bifurcations in a delayed predator–prey diffusion system with a functional response

The present paper is concerned with a delayed predator–prey diffusion system with a Beddington–DeAngelis functional response and homogeneous Neumann boundary conditions. If the positive constant steady state of the corresponding system without delay is stable, by choosing the delay as the bifurcation parameter, we can show that the increase of the delay can not only cause spatially homogeneous Hopf bifurcation at the positive constant steady state but also give rise to spatially heterogeneous ones. In particular, under appropriate conditions, we find that the system has a Bogdanov–Takens singularity at the positive constant steady state, whereas this singularity does not occur for the corresponding system without diffusion. In addition, by applying the normal form theory and center manifold theorem for partial functional differential equations, we give normal forms of Hopf bifurcation and Bogdanov–Takens bifurcation and the explicit formula for determining the properties of spatial Hopf bifurcations.     

Global stability and Hopf bifurcation of a plankton model with time delay

A differential delay equation model with a discrete time delay and a distributed time delay is introduced to simulate zooplankton–nutrient interaction. The differential inequalities’ methods and standard Hopf bifurcation analysis are applied. Some sufficient conditions are obtained for persistence and for the global stability of the unique positive steady state, respectively. It was shown that there is a Hopf bifurcation in the model by using the discrete time delay as a bifurcation parameter.     

Bifurcation analysis of the Gierer–Meinhardt system with a saturation in the activator production

The reaction–diffusion Gierer–Meinhardt system with a saturation in the activator production is considered. Stability of the unique positive constant steady state solution is analysed, and associated Hopf bifurcations and steady state bifurcations are obtained. A global bifurcation diagram of non-trivial periodic orbits and steady state solutions with respect to key system parameters is obtained, which improves the understanding of dynamics of Gierer–Meinhardt system with a saturation in different parameter regimes.     

Dynamics of a competitive Lotka-Volterra system with three delays

In this paper, a competitive Lotka-Volterra system with three delays is investigated. By choosing the sum τ of three delays as a bifurcation parameter, we show that in the above system, Hopf bifurcation at the positive equilibrium can occur as τ crosses some critical values. And we obtain the formulae determining direction of Hopf bifurcation and stability of the bifurcating periodic solutions by using the normal form theory and center manifold theorem. Finally, numerical simulations supporting our theoretical results are also included.     

Detection of Hopf points by counting sectors in the complex plane

Summary. Let ( real) be a family of real by matrices. A value of is called a Hopf value if has a conjugate pair of purely imaginary eigenvalues , . We describe a technique for detecting Hopf values based on the evolution of the Schur complement of in a bordered extension of where varies along the positive imaginary axis of the complex plane. We compare the efficiency of this method with more obvious methods such as the use of the QR algorithm and of the determinant function of as well as with recent work on the Cayley transform. In particular, we show the advantages of the Schur complement method in the case of large sparse matrices arising in dynamical problems by discretizing boundary value problems. The Hopf values of the Jacobian matrices are important in this setting because they are related to the Hopf bifurcation phenomenon where steady state solutions bifurcate into periodic solutions. Received September 15, 1994 / Revised version received July 7, 1995     

一类带无选择性捕获和时滞的捕食食饵系统的Hopf分支分析

本文主要研究了一个改进的带时滞和无选择捕获函数的捕食-食饵生态经济系统的稳定性和Hopf分支.利用微分代数系统的稳定性理论和分支理论,得到了系统正平衡点稳定性的条件,以及当时滞τ作为分支参数时系统产生Hopf分支的条件.对Leslie-Gower捕食-食饵模型进行了一定程度的完善,使得建立的模型更符合实际情况,因此得到的结论也更加科学.     

Bifurcation analysis of a modified Holling-Tanner predator-prey model with time delay

In this paper, a modified Holling-Tanner predator-prey model with time delay is considered. By regarding the delay as the bifurcation parameter, the local asymptotic stability of the positive equilibrium is investigated. Meanwhile, we find that the system can also undergo a Hopf bifurcation of nonconstant periodic solution at the positive equilibrium when the delay crosses through a sequence of critical values. In particular, we study the direction of Hopf bifurcation and the stability of bifurcated periodic solutions, an explicit algorithm is given by applying the normal form theory and the center manifold reduction for functional differential equations. Finally, numerical simulations supporting the theoretical analysis are also included.     

Bifurcation and spatiotemporal patterns in a homogeneous diffusive predator-prey system

A diffusive predator-prey system with Holling type-II predator functional response subject to Neumann boundary conditions is considered. Hopf and steady state bifurcation analysis are carried out in details. In particular we show the existence of multiple spatially non-homogeneous periodic orbits while the system parameters are all spatially homogeneous. Our results and global bifurcation theory also suggest the existence of loops of spatially non-homogeneous periodic orbits and steady state solutions. These results provide theoretical evidences to the complex spatiotemporal dynamics found by numerical simulation.     

Stability and Hopf bifurcation analysis in a novel congestion control model with communication delay

In this paper, we investigated Hopf bifurcation by analyzing the distributed ranges of eigenvalues of characteristic linearized equation. Using communication delay as the bifurcation parameter, linear stability criteria dependent on communication delay have also been derived, and, furthermore, the direction of Hopf bifurcation as well as stability of periodic solution for the exponential RED algorithm with communication delay is studied. We find that the Hopf bifurcation occurs when the communication delay passes a sequence of critical values. The stability and direction of the Hopf bifurcation are determined by applying the normal form theory and the center manifold theorem. Finally, a numerical simulation is presented to verify the theoretical results.     

Hopf Bifurcation for a Susceptible-Infective Model with Infection-Age Structure

An SIS model is investigated in which the infective individuals are assumed to have an infection-age structure. The model is formulated as an abstract non-densely defined Cauchy problem. We study some dynamical properties of the model by using the theory of integrated semigroups, the Hopf bifurcation theory and the normal form theory for semilinear equations with non-dense domain. Qualitative analysis indicates that there exist some parameter values such that this SIS model has a non-trivial periodic solution which bifurcates from the positive equilibrium. Furthermore, the explicit formulae are given to determine the direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions. Numerical simulations are also carried out to support our theoretical results.