Bifurcations in a predator–prey model in patchy environment with diffusion

In this paper we formulate a predator–prey system in two patches in which the per capita migration rate of each species is influenced only by its own density, i.e. there is no response to the density of the other one. Numerical studies show that at a critical value of the bifurcation parameter the system undergoes a Turing bifurcation, i.e. the stable constant steady state loses its stability and spatially non-constant stationary solutions, a pattern emerge.     

Stationary distribution and extinction of a stochastic predator–prey model with distributed delay

In this paper, we focus on a stochastic predator–prey model with distributed delay. We first obtain the existence of a stationary distribution to the positive solutions by stochastic Lyapunov function method. Then we establish sufficient conditions for extinction of the predator population, that is, the prey population is survival and the predator population is extinct.     

Periodic solutions of “predator–prey” systems with continuous delay and periodic coefficients

We prove the existence of positive ω-periodic solutions for some “predator–prey” systems with continuous delay of the argument for the case where the parameters of these systems are specified by ω-periodic continuous positive functions.     

Permanence of periodic predator–prey system with two predators and stage structure for prey

A stage-structured three-species predator–prey system with Beddington–DeAngelis and Holling IV functional response is proposed and analyzed. Based on the comparison theorem, some sufficient and necessary conditions are derived for permanence of the system. Finally, two examples are presented to illustrate the application of our main results.     

Permanence and periodic solutions for a diffusive ratio-dependent predator–prey system

This paper considers a diffusive predator–prey model, in which there is a ratio-dependent functional response with Holling III type. We establish some sufficient conditions for the ultimate boundedness of solutions and permanence of this system. The existence of a unique globally stable periodic solution is also presented.     

Dynamics in a diffusive predator–prey system with a constant prey refuge and delay

In this paper, a diffusive predator–prey system with a constant prey refuge and time delay subject to Neumann boundary condition is considered. Local stability and Turing instability of the positive equilibrium are studied. The effect of time delay on the model is also obtained, including locally asymptotical stability and existence of Hopf bifurcation at the positive equilibrium. And the properties of Hopf bifurcation are determined by center manifold theorem and normal form theorem of partial functional differential equations. Some numerical simulations are carried out.     

Hopf bifurcations in a predator–prey system with a discrete delay and a distributed delay

A delayed Lotka–Volterra two-species predator–prey system with discrete hunting delay and distributed maturation delay for the predator population described by an integral with a strong delay kernel is considered. By linearizing the system at the positive equilibrium and analyzing the associated characteristic equation, the asymptotic stability of the positive equilibrium is investigated and Hopf bifurcations are demonstrated. It is found that under suitable conditions on the parameters the positive equilibrium is asymptotically stable when the hunting delay is less than a certain critical value and unstable when the hunting delay is greater than this critical value. Meanwhile, according to the Hopf bifurcation theorem for functional differential equations (FDEs), we find that the system can also undergo a Hopf bifurcation of nonconstant periodic solution at the positive equilibrium when the hunting delay crosses through a sequence of critical values. In particular, by applying the normal form theory and the center manifold reduction for FDEs, an explicit algorithm determining the direction of Hopf bifurcations and the stability of bifurcating periodic solutions occurring through Hopf bifurcations is given. Finally, to verify our theoretical predictions, some numerical simulations are also included at the end of this paper.     

Permanence and extinction in competitive Lotka–Volterra systems with delays

In this paper, a class of competitive Lotka–Volterra systems are considered that have distributed delays and constant coefficients on interaction terms and have time dependent growth rate vectors with an asymptotic average. Under the assumption that all proper subsystems are permanent, it is shown that the asymptotic behaviour of the system is determined by the relationship between an equilibrium and a nullcline plane of the corresponding autonomous system: if the equilibrium is below the plane then the system is permanent; if the equilibrium is above the plane then this species will go extinct in an exponential rate while the other species will survive. Similar asymptotic behaviour is also retained under an alternative assumption.     

Modelling a predator–prey system with infected prey in polluted environment

A predator–prey model with logistic growth in prey is modified by introducing an SIS parasite infection in the prey. We have studied the combined effect of environmental toxicant and disease on prey–predator system. It is assumed in this paper that the environmental toxicant affects both prey and predator population and the infected prey is assumed to be more vulnerable to the toxicant and predation compared to the sound prey individuals. Thresholds are identified which determine when system persists and disease remains endemic.     

Bifurcation and chaos in a ratio-dependent predator–prey system with time delay

In this paper, a ratio-dependent predator–prey model with time delay is investigated. We first consider the local stability of a positive equilibrium and the existence of Hopf bifurcations. By using the normal form theory and center manifold reduction, we derive explicit formulae which determine the stability, direction and other properties of bifurcating periodic solutions. Finally, we consider the effect of impulses on the dynamics of the above time-delayed population model. Numerical simulations show that the system with constant periodic impulsive perturbations admits rich complex dynamic, such as periodic doubling cascade and chaos.     

Stability and Hopf bifurcation in a diffusive predator–prey system with delay effect

This paper is concerned with a delayed predator–prey system with diffusion effect. First, the stability of the positive equilibrium and the existence of spatially homogeneous and spatially inhomogeneous periodic solutions are investigated by analyzing the distribution of the eigenvalues. Next the direction and the stability of Hopf bifurcation are determined by the normal form theory and the center manifold reduction for partial functional differential equations. Finally, some numerical simulations are carried out for illustrating the theoretical results.     

Bifurcation and control of a bioeconomic model of a prey–predator system with a time delay

In this paper, we analyze the dynamical behaviour of a bioeconomic model system using differential algebraic equations. The system describes a prey–predator fishery with prey dispersal in a two-patch environment, one of which is a free fishing zone and other is a protected zone. It is observed that a singularity-induced bifurcation phenomenon appears when a variation of the economic interest of harvesting is taken into account. We have incorporated a state feedback controller to stabilize the model system in the case of positive economic interest. A discrete-type gestational delay of predators is incorporated, and its effect on the dynamical behaviour of the model is analyzed. The occurrence of Hopf bifurcation of the proposed model with positive economic profit is shown in the neighbourhood of the coexisting equilibrium point through considering the delay as a bifurcation parameter. Finally, some numerical simulations are given to verify the analytical results, and the system is analyzed through graphical illustrations.     

A predator–prey system with stage-structure for predator and nonlocal delay

This paper deals with the behavior of solutions to the reaction–diffusion system under homogeneous Neumann boundary condition, which describes a prey–predator model with nonlocal delay. Sufficient conditions for the global stability of each equilibrium are derived by the Lyapunov functional and the results show that the introduction of stage-structure into predator positively affects the coexistence of prey and predator. Numerical simulations are performed to illustrate the results.     

Global dynamics of a predator–prey model with time delay and stage structure for the prey

A stage-structured predator–prey system with Holling type-II functional response and time delay due to the gestation of predator is investigated. By analyzing the characteristic equations, the local stability of each of feasible equilibria of the system is discussed and the existence of a Hopf bifurcation at the coexistence equilibrium is established. By means of the persistence theory on infinite dimensional systems, it is proven that the system is permanent if the coexistence equilibrium exists. By using Lyapunov functionals and LaSalle invariant principle, it is shown that the trivial equilibrium is globally stable when both the predator-extinction equilibrium and the coexistence equilibrium are not feasible, and that the predator-extinction equilibrium is globally asymptotically stable if the coexistence equilibrium does not exist, and sufficient conditions are derived for the global stability of the coexistence equilibrium. Numerical simulations are carried out to illustrate the main theoretical results.     

Multiple periodic solutions in generalized Gause-type predator–prey systems with non-monotonic numerical responses

By using a continuation theorem based on coincidence degree theory, we establish easily verifiable criteria for the existence of multiple positive periodic solutions in periodic Gause-type predator–prey systems with non-monotonic numerical responses and time delays. As corollaries, some applications are listed. In particular, our results improve and supplement those obtained by Chen [Y. Chen, Multiple periodic solutions of delayed predator–prey systems with type IV functional responses, Nonlinear Anal. Real World Appl. 5 (2004) 45–53].     

Modeling and analysis of a prey–predator system with disease in the prey

A three dimensional ecoepidemiological model consisting of susceptible prey, infected prey and predator is proposed and analysed in the present work. The parameter delay is introduced in the model system for considering the time taken by a susceptible prey to become infected. Mathematically we analyze the dynamics of the system such as, boundedness of the solutions, existence of non-negative equilibria, local and global stability of interior equilibrium point. Next we choose delay as a bifurcation parameter to examine the existence of the Hopf bifurcation of the system around its interior equilibrium. Moreover we use the normal form method and center manifold theorem to investigate the direction of the Hopf bifurcation and stability of the bifurcating limit cycle. Some numerical simulations are carried out to support the analytical results.     

Multiple periodic solutions of delayed predator–prey systems with type IV functional responses

In this paper, we considered a periodic predator–prey system with a type IV functional response, which incorporates the periodicity of the environment. Sufficient conditions for the existence of multiple positive periodic solutions are established by applying the continuation theorem. This is the first time that multiple periodic solutions are obtained by using the theory of coincidence degree. Moreover, unlike other types of functional responses, a type IV functional response declines at high prey densities. Thus the existing arguments for obtaining bounds of solutions to the operator equation Lx=λNx are inapplicable to our case and some new arguments are employed for the first time.     

Evolution in predator–prey systems

We study the adaptive dynamics of predator–prey systems modeled by a dynamical system in which the traits of predators and prey are allowed to evolve by small mutations. When only the prey are allowed to evolve, and the size of the mutational change tends to 0, the system does not exhibit long term prey coexistence and the trait of the resident prey type converges to the solution of an ODE. When only the predators are allowed to evolve, coexistence of predators occurs. In this case, depending on the parameters being varied, we see that (i) the number of coexisting predators remains tight and the differences in traits from a reference species converge in distribution to a limit, or (ii) the number of coexisting predators tends to infinity, and we calculate the asymptotic rate at which the traits of the least and most “fit” predators in the population increase. This last result is obtained by comparison with a branching random walk killed to the left of a linear boundary and a finite branching–selection particle system.