Optimal harvesting policies for periodic Gompertz systems

In this paper, we study the periodic Gompertz system with harvesting. First, we analyze the system with continuous harvesting and obtain the maximum annual-sustainable yield, the optimal harvesting effort and the optimal population level for such a system. Then, the harvesting is assumed to occur at fixed moments every year, and we establish the Gompertz system with impulsive perturbation. And we investigate the impulsive harvesting policy to maximize the annual yield and to keep the population sustainable development. At last, the optimal results of the impulsive harvesting system are compared with those of the continuous harvesting system.     

Optimal pulse fishing policy in stage-structured models with birth pulses

In this paper, we propose exploited models with stage structure for the dynamics in a fish population for which periodic birth pulse and pulse fishing occur at different fixed time. Using the stroboscopic map, we obtain an exact cycle of system, and obtain the threshold conditions for its stability. Bifurcation diagrams are constructed with the birth rate (or pulse fishing time or harvesting effort) as the bifurcation parameter, and these are observed to display complex dynamic behaviors, including chaotic bands with period windows, period-doubling, multi-period-halving and incomplete period-doubling bifurcation, pitch-fork and tangent bifurcation, non-unique dynamics (meaning that several attractors or attractor and chaos coexist) and attractor crisis. This suggests that birth pulse and pulse fishing provide a natural period or cyclicity that make the dynamical behaviors more complex. Moreover, we show that the pulse fishing has a strong impact on the persistence of the fish population, on the volume of mature fish stock and on the maximum annual-sustainable yield. An interesting result is obtained that, after the birth pulse, the population can sustain much higher harvesting effort if the mature fish is removed as early as possible.     

Optimal harvesting policy for stochastic Logistic population model

In this paper, we consider some optimal harvesting policies for single population models, in which the harvest effort and the intrinsic growth rate are disturbed by environment noises. We choose the maximum sustainable yield and the maximum retained profits as two management objectives, and obtain the optimal harvesting policies, respectively. For the two objectives, we give the optimal harvest effort that maximizes the sustainable yield (or retained profits), the maximum of expectation of sustainable yield (or retained profits) and the corresponding variance. Their explicit expressions are determined by the coefficients of equation and the disturbance intensity.     

周期Gompertz生态系统中的最优脉冲控制收获策略

以周期Gompertz系统为基础,讨论了周期变化的单种群生物资源的收获优化问题及种群的动力学性质.在单位收获努力量假设下,以最大可持续收获量为管理目标,确定了线性收获下的最优收获策略,获得了最优收获努力量、最大可持续收获及相应的最优种群水平的显示表达式,为自然资源的开发和利用提供了理论依据.     

Asymptotically periodic solution and optimal harvesting policy for Gompertz system

In this paper, the single species modelled by (asymptotically) periodic Gompertz equation is investigated. It is shown that the (asymptotically) periodic system has a unique (asymptotically) periodic solution which is globally asymptotically stable for the positive solution. When the nonautonomous Gompertz equation is subject to harvesting, we study the optimal harvesting policy for the periodic system and obtain the corresponding optimal population level and the maximum sustainable yield. Further, when the functions in the exploited Gompertz system are stably bounded functions, we study the ultimately optimal harvesting policy. By choosing the average limiting maximum sustainable yield as management objective, the corresponding optimal population level is determined.     

广义Richards-Gilpin-Ayala模型的捕获优化问题

在Richards-Gilpin-Ayala模型的基础上,提出了一类更广泛的数学模型—广义Richards-Gilpin-Ayala模型.进而讨论该模型单种群生物资源的捕获优化问题,分析了被开发生物种群的动力学性质.在单位捕获努力量假定下,以最大可持续捕获量为管理目标,确定了线性捕获下的最优捕获策略,得到了最优捕获努力量,最大可持续收获及相应的最优种群水平的显式表达式.这些结果推广了相关文献中关于Schaefer模型、广义Logistic模型的相应结果.     

Optimal impulsive harvesting on non-autonomous Beverton–Holt difference equations

Many recent advances in the theory of the optimal economic exploitation of renewable fish resources have been gained by applying optimal control theory. However, despite these successes, much less is known about how seasonal environments affect the maximum sustainable yield (MSY) (or population persistence) and any effects of relations between intensity and frequency of harvesting. Assuming that fish populations follow Beverton–Holt equations we investigated impulsive harvesting in seasonal environments, focusing on both economic aspects and resource sustainability. We first investigated the existence and stability of a periodic solution and its analytic formula, and then showed that the population persistence depends on the intensity and frequency of harvesting. With the MSY as a management objective, we investigated optimal impulsive harvesting policies. The optimal harvesting effort that maximizes the sustainable yield, the corresponding optimal population level, and the MSY are obtained by using discrete Euler–Lagrange equations and product formulae, and their explicit expressions were obtained in terms of the intrinsic growth rate, the carrying capacity, and the impulsive moments. These results imply that harvest timing is of crucial importance to the MSY. Since impulsive differential equations incorporate elements of continuous and discrete systems, we can apply all results obtained for Beverton–Holt equations with impulsive effects to periodic logistic equations with impulsive harvesting.     

Optimal impulsive harvesting on non-autonomous Beverton–Holt difference equations

Many recent advances in the theory of the optimal economic exploitation of renewable fish resources have been gained by applying optimal control theory. However, despite these successes, much less is known about how seasonal environments affect the maximum sustainable yield (MSY) (or population persistence) and any effects of relations between intensity and frequency of harvesting. Assuming that fish populations follow Beverton–Holt equations we investigated impulsive harvesting in seasonal environments, focusing on both economic aspects and resource sustainability. We first investigated the existence and stability of a periodic solution and its analytic formula, and then showed that the population persistence depends on the intensity and frequency of harvesting. With the MSY as a management objective, we investigated optimal impulsive harvesting policies. The optimal harvesting effort that maximizes the sustainable yield, the corresponding optimal population level, and the MSY are obtained by using discrete Euler–Lagrange equations and product formulae, and their explicit expressions were obtained in terms of the intrinsic growth rate, the carrying capacity, and the impulsive moments. These results imply that harvest timing is of crucial importance to the MSY. Since impulsive differential equations incorporate elements of continuous and discrete systems, we can apply all results obtained for Beverton–Holt equations with impulsive effects to periodic logistic equations with impulsive harvesting.     

Optimal impulsive harvesting policy for single population

In this paper, we established the exploitation of impulsive harvesting single autonomous population model by Logistic equation. By some special methods, we analysis the impulsive harvesting population equation and obtain existence, the explicit expression and global attractiveness of impulsive periodic solutions for constant yield harvest and proportional harvest. Then, we choose the maximum sustainable yield as management objective, and investigate the optimal impulsive harvesting policies respectively. The optimal harvest effort that maximizes the sustainable yield per unit time, the corresponding optimal population levels are determined. At last, we point out that the continuous harvesting policy is superior to the impulsive harvesting policy, however, the latter is more beneficial in realistic operation.     

Optimal harvesting policy for general stochastic Logistic population model

We consider some optimal harvesting policies for a general stochastic Logistic population model. For two management objectives, that are maximum sustainable yield and the maximum retained profits, the optimal harvesting policies are obtained. Meanwhile, the optimal harvest effort, the maximum of expectation of sustainable yield (or retained profits) and the corresponding variance are given.     

周期性Gompertz差分模型的最优脉冲收获策略

考虑一个具有周期性脉冲收获的Gompertz差分系统,推导了保证种群系统持续生存、绝灭以及存在全局吸引的正脉冲周期解的充要条件,以一个周期内持续产量最大化为管理目标,通过利用离散的Pontryagin最大值原理获得了最优的脉冲收获策略,推广了现有的结论.     

POPULATION AND HARVESTING THEORY FOR NONLINEAR SEX AND AGE-STRUCTURED RESOURCES

A sex-age-structured population model with density dependence in the conversion of reproductive potentials into zygotes and in first year survivorship is described. The model has two equilibria; the smallest is mathematically unstable, and the origin and the larger equilibrium are locally stable. The population can thus go extinct for certain initial states, or if the two equilibria coincide. The ratio between the two equilibria can be regarded as a measure of the risk of extinction, since it is related to the chance that detrimental environmental conditions will cause the population to enter the region of attraction of the origin. In simple monoecious models, recovery to former levels is only possible provided that the population is not driven to extinction before harvesting effort is reduced. Ratios between the two unexploited equilibria, and between the stable unexploited equilibrium and the recruitment level at which the two equilibria coincide are given solely in terms of the degree of density dependence in the model. I show that the harvesting strategy which maximizes the equilibrium yield has a four age form, involving harvesting of at most two male and two female age classes. Out of ten commercial Pacific groundfish species, knife-edge selectivity sustainable yields of eight are at least 90% of ultimate sustainable yield (USY). With no effort restrictions, the range of lengths at first capture which achieve more than 60% of USY is narrow. When one of the sexes is not harvested, sustainable yield is between 20% and 80% of USY, but lowest when females are not harvested.     

OPTIMAL HARVESTING POLICY FOR INSHORE-OFFSHORE FISHERY MODEL WITH IMPULSIVE DIFFUSION

This article studies the inshore-offshore fishery model with impulsive diffusion. The existence and global asymptotic stability of both the trivial periodic solution and the positive periodic solution are obtained. The complexity of this system is also analyzed. Moreover, the optimal harvesting policy are given for the inshore subpopulation, which includes the maximum sustainable yield and the corresponding harvesting effort.     

Optimal harvesting policy for a stochastic predator–prey model

Optimal harvesting of a stochastic predator–prey model is considered in this paper. Sufficient and necessary criteria for the existence of optimal harvesting strategy are obtained. At the same time, the optimal harvest effort and the maximum of sustainable yield are given.     

SUSTAINABLE YIELDS IN FISHERIES: UNCERTAINTY,RISK‐AVERSION,AND MEAN‐VARIANCE ANALYSIS

Abstract We consider a model of a fishery in which the dynamics of the unharvested fish population are given by the stochastic logistic growth equation Similar to the classical deterministic analogon, we assume that the fishery harvests the fish population following a constant effort strategy. In the first step, we derive the effort level that leads to maximum expected sustainable yield, which is understood as the expectation of the equilibrium distribution of the stochastic dynamics. This replaces the nonzero fixed point in the classical deterministic setup. In the second step, we assume that the fishery is risk averse and that there is a tradeoff between expected sustainable yield and uncertainty measured in terms of the variance of the equilibrium distribution. We derive the optimal constant effort harvesting strategy for this problem. In the final step, we consider an approach that we call the mean‐variance analysis to sustainable fisheries. Similar as in the now classical mean‐variance analysis in finance, going back to Markowitz [1952] , we study the problem of maximizing expected sustainable yields under variance constraints, and with this, minimizing the variance, e.g., risk, under guaranteed minimum expected sustainable yields. We derive explicit formulas for the optimal fishing effort in all four problems considered and study the effects of uncertainty, risk aversion, and mean reversion speed on fishing efforts.     

Optimal Harvesting for a Stochastic Predator-prey Model with S-type Distributed Time Delays

In this paper, the optimization problem of harvesting for a stochastic predator-prey model with S-type distributed time delays (which contain both discrete time delays and continuously distributed time delays) is studied by using ergodic method. Sufficient and necessary conditions for the existence of optimal harvesting strategy are obtained. Moreover, the optimal harvesting effort (OHE, for short) and the maximum of expectation of sustainable yield (MESY, for short) are given. Some numerical simulations are introduced to illustrate our main results.     

HARVESTING STRATEGIES FOR A MIGRATORY FISH STOCK BASED ON A MODEL WITH UNCERTAINTY

We model a species of fish with breeding ground protected, and harvesting is carried out in an adjacent area. There is migration of fish from the breeding ground to the adjacent area. We demonstrate that in the presence of bounded uncertain disturbances, the strategy of harvesting the fish biomass at the maximum sustainable yield level is indeed optimal in the long term consideration.     

Optimal and feedback strategies for managing multicohort populations

A Beverton and Holt type linear cohort dynamics model is integrated and combined with a nonlinear stock-recruitment relationship to obtain a discrete-time multicohort harvesting model. Assuming that each age class is individually controllable, it is shown, subject to certain assumptions, that the optimal harvesting strategy is to drive the population to the maximum sustainable yield solution in one time step. In most fisheries, this controllability assumption is not met and harvesting is agewise nonselective. In this case, it may be preferable to implement a harvesting policy based on suboptimal constant effort or stock level feedback strategies, rather than implement a more complicated optimal policy. This question is addressed through numerical studies on the management of an anchovy fishery.Dedicated to G. LeitmannThe author would like to thank M. Mangel, W. Reed, P. Sullivan, and G. Swartzman for commenting on a draft of this paper.     

Global stability in a structured population competition model with distributed maturation delay and harvesting

This paper is concerned with a delay differential equation model for the interaction between two species, the adult members of which are in competition, with stage-structure and harvesting of the mature and immature members of each species. The maturation delay for each species is modelled as a distribution, to allow for the possibility that individuals may take a different amount of time to mature. General birth and death rate functions are used. We find that the dynamics of the model depends largely on the birth and death functions, which depend on the total number of adults.We study the dynamics of our model analytically and we present results on the positivity and boundedness of the solutions, and global stability results are established for each equilibrium.     

Qualitative analysis and applications of a kind of state-dependent impulsive differential equations

This paper analyzes a certain type of impulsive differential equations (IDEs). Several useful theorems for its periodic solutions and their stabilities are given. The key idea is that a periodically time-dependent IDE can be transformed into the state-dependent IDE. As applications of our theory, the optimization problems in population dynamics are studied. That is, the maximum sustainable yields of single population models with periodically impulsive constant harvesting are discussed. Furthermore, we apply these results to the studies of the order-1 periodic solutions and their stability of a single population model with stage structure in which the mature is impulsively proportionally harvested while the immature is impulsively added with the constant.