Permanence and extinction of a three-species ratio-dependent food chain model with delay and prey diffusion

In this paper, we study a diffusive three-species ratio-dependent food chain model, using differential inequality, to obtain sufficient conditions that ensure the permanence of the system and the extinction of predator species. Our results reinforce the main result of Sun Wen, Shihua Chen and Huihai Mei [Positive periodic solution of a more realistic three-species Lotka-Volterra model with delay and density regulation, Chaos, Solitons and Fractals, in press].     

Global attractivity and permanence of a SVEIR epidemic model with pulse vaccination and time delay

In this study, we propose a new SVEIR epidemic disease model with time delay, and analyze the dynamic behavior of the model under pulse vaccination. Pulse vaccination is an effective strategy for the elimination of infectious disease. Using the discrete dynamical system determined by the stroboscopic map, we obtain an ‘infection-free’ periodic solution. We also show that the ‘infection-free’ periodic solution is globally attractive when some parameters of the model under appropriate conditions. The permanence of the model is investigated analytically. Our results indicate that a large vaccination rate or a short pulse of vaccination or a long latent period is a sufficient condition for the extinction of the disease.     

Dynamic behaviors of a delay differential equation model of plankton allelopathy

In this paper, we consider a modified delay differential equation model of the growth of n-species of plankton having competitive and allelopathic effects on each other. We first obtain the sufficient conditions which guarantee the permanence of the system. As a corollary, for periodic case, we obtain a set of delay-dependent condition which ensures the existence of at least one positive periodic solution of the system. After that, by means of a suitable Lyapunov functional, sufficient conditions are derived for the global attractivity of the system. For the two-dimensional case, under some suitable assumptions, we prove that one of the components will be driven to extinction while the other will stabilize at a certain solution of a logistic equation. Examples show the feasibility of the main results.     

Stochastic modeling of phytoplankton allelopathy

The study of dynamic interactions between two competing phytoplankton species in the presence of toxic substances is an active field of research due to the global increase of harmful phytoplankton blooms. Ordinary differential equation models for two competing phytoplankton species, when one or both the species liberate toxic substances, are unable to capture the oscillatory and highly variable growth of phytoplankton populations. The deterministic formulation never predicts the sudden localized extinction of certain species. These obstacles of mathematical modeling can be overcome if we include stochastic variability in our modeling approach. In this investigation, we construct stochastic models of allelopathic interactions between two competing phytoplankton species as a continuous time Markov chain model as well as an Itô stochastic differential equation model. Approximate extinction probabilities for both species are obtained analytically for the continuous time Markov chain model. Analytical estimates are validated with the help of numerical simulations.     

Dynamics of species in a model with two predators and one prey

In this paper, we study a predator-prey model which has one prey and two predators with Beddington-DeAngelis functional responses. Firstly, we establish a set of sufficient conditions for the permanence and extinction of species. Secondly, the periodicity of positive solutions is studied. Thirdly, by using Liapunov functions and the continuation theorem in coincidence degree theory, we show the global asymptotic stability of such solutions. Finally, we give some numerical examples to illustrate the behavior of the model.     

Dynamics of a neutral delay equation for an insect population with long larval and short adult phases

We present a global study on the stability of the equilibria in a nonlinear autonomous neutral delay differential population model formulated by Bocharov and Hadeler. This model may be suitable for describing the intriguing dynamics of an insect population with long larval and short adult phases such as the periodical cicada. We circumvent the usual difficulties associated with the study of the stability of a nonlinear neutral delay differential model by transforming it to an appropriate non-neutral nonautonomous delay differential equation with unbounded delay. In the case that no juveniles give birth, we establish the positivity and boundedness of solutions by ad hoc methods and global stability of the extinction and positive equilibria by the method of iteration. We also show that if the time adjusted instantaneous birth rate at the time of maturation is greater than 1, then the population will grow without bound, regardless of the population death process.     

EXTINCTION IN A GILPIN-AYALA COMPETITION SYSTEM WITH THE EFFECT OF TOXIC SUBSTANCES

In this paper,we consider a two dimensional Gilpin-Ayala competition system with the effect of toxic substances.We prove that one of components is driven to extinction while the other one is stable under some conditions.As a result,we generalize the previous results.     

Existence of multiple periodic solutions for an SIR model with seasonality

We study an SIR model with a seasonal contact rate and a staged treatment strategy, which is an extension of our previous work [Z. Bai, Y. Zhou, Existence of two periodic solutions for a non-autonomous SIR epidemic model, Appl. Math. Model. 35 (2011) 382-391]. It is proved that the persistence and extinction of the disease are determined by the basic reproductive number (R₀) and a threshold parameter (R_c). It is obtained that the model exhibits two different bistable behaviors under certain conditions: the stable disease-free state coexists with a stable endemic periodic solution, and three endemic periodic solutions coexist with two of them being stable. Numerical simulations are presented to illustrate theoretical results.     

Diffusive monotonicity and threshold dynamics of delayed reaction diffusion equations

For abstract functional differential equations and reaction-diffusion equations with delay, an exponential ordering is introduced which takes into account the spatial diffusion. The induced monotonicity of the solution semiflows is established and applied to describe the threshold dynamics (extinction or persistence/convergence to positive equilibria) for a nonlocal and delayed reaction-diffusion population model.     

On the behavior of the Lorenz equation backward in time

The sets of solutions to the Lorenz equations that exist backward in time and are bounded at an exponential rate determined by the eigenvalues of the linear part of the equation are examined. The set associated with the middle eigenvalue is shown to project surjectively onto a plane, thereby providing a lower estimate for its dimension. Specific bounds are also found for a cone containing this set.     

Robustness of nonuniform exponential dichotomies in Banach spaces

We give conditions for the robustness of nonuniform exponential dichotomies in Banach spaces, in the sense that the existence of an exponential dichotomy for a given linear equation x^′=A(t)x persists under a sufficiently small linear perturbation. We also establish the continuous dependence with the perturbation of the constants in the notion of dichotomy and of the “angles” between the stable and unstable subspaces. Our proofs exhibit (implicitly) the exponential dichotomies of the perturbed equations in terms of fixed points of appropriate contractions. We emphasize that we do not need the notion of admissibility (of bounded nonlinear perturbations). We also obtain related robustness results in the case of nonuniform exponential contractions. In addition, we establish an appropriate version of robustness for nonautonomous dynamical systems with discrete time.     

The linear and nonlinear diffusion of the competitive Lotka–Volterra model

This paper has studied the effects of linear and nonlinear diffusion of the competitive Lotka–Volterra model, and has investigated how the linear and nonlinear diffusions lead from the extinction of one species to the persistence or global asymptotic stability of all species. This research has important implications in the design of nature reserves.     

On nonuniform exponential dichotomy of evolution operators in Banach spaces

The problem of nonuniform exponential dichotomy of evolution operators in Banach spaces is considered. Connections between this concept and admissibility of the pair (C ₀,C ₀) are established. Generalizations to the nonuniform case of some results of Van Min, Räbiger and Schnaubelt ([MRS]) are obtained. It is shown that an implication from the uniform case is not true for nonuniform exponential dichotomy. The results are applicable for general time-varying linear equations with unbounded coefficients in Banach spaces.     

Short-term recurrent chaos and role of Toxin Producing Phytoplankton (TPP) on chaotic dynamics in aquatic systems

We propose a new mathematical model for aquatic populations. This model incorporates mutual interference in all the three populations and an extra mortality term in zooplankton population and also taking into account the toxin liberation process of TPP population. The proposed model generalizes several other known models in the literature. The principal interest in this paper is in a numerical study of the model’s behaviour. It is observed that both types of food chains display same type of chaotic behaviour, short-term recurrent chaos, with different generating mechanisms. Toxin producing phytoplankton (TPP) reduces the grazing pressure of zooplankton. To observe the role of TPP, we consider Holling types I, II and III functional forms for this process. Our study suggests that toxic substances released by TPP population may act as bio-control by changing the state of chaos to order and extinction.     

Differential equations with bounded positive Green’s functions and generalized Aizerman’s hypothesis

We derive conditions for the positivity and boundedness of the Green functions of the higher order linear nonautonomous ODE. By virtue of these conditions, the existence of positive solutions for a class of nonlinear equations is proved. In addition, upper and lower estimates for the Green functions are established. Moreover, it is shown that nonlinear equations, having separated nonautonomous linear parts, satisfy the generalized Aizerman hypothesis on absolute stability, if they have the positive Green functions.     

Qualitative analysis and applications of a kind of state-dependent impulsive differential equations

This paper analyzes a certain type of impulsive differential equations (IDEs). Several useful theorems for its periodic solutions and their stabilities are given. The key idea is that a periodically time-dependent IDE can be transformed into the state-dependent IDE. As applications of our theory, the optimization problems in population dynamics are studied. That is, the maximum sustainable yields of single population models with periodically impulsive constant harvesting are discussed. Furthermore, we apply these results to the studies of the order-1 periodic solutions and their stability of a single population model with stage structure in which the mature is impulsively proportionally harvested while the immature is impulsively added with the constant.     

Optimal regularity of robustness for parameterized perturbations

For exponential dichotomies defined by nonautonomous linear equations, we show that sufficiently small C¹-parameterized perturbations originate a family of exponential dichotomies of class C¹ in the parameter. We consider the general case of nonuniform exponential dichotomies, and also the general case of arbitrary growth rates of the form eλρ(t) where ρ is an arbitrary function. This includes the usual exponential behavior as a very special case when ρ(t)=t.     

Lyapunov regularity of impulsive differential equations

For linear impulsive differential equations, we give a simple criterion for the existence of a nonuniform exponential dichotomy, which includes uniform exponential dichotomies as a very special case. For this we introduce the notion of Lyapunov regularity for a linear impulsive differential equation, in terms of the so-called regularity coefficient. The theory is then used to show that if the Lyapunov exponents are nonzero, then there is a nonuniform exponential behavior, which can be expressed in terms of the Lyapunov exponents of the differential equation and of the regularity coefficient. We also consider the particular case of nonuniform exponential contractions when there are only negative Lyapunov exponents. Having this relation in mind, it is also of interest to provide alternative characterizations of Lyapunov regularity, and particularly to obtain sharp lower and upper bound for the regularity coefficient. In particular, we obtain bounds expressed in terms of the matrices defining the impulsive linear system, and we obtain characterizations in terms of the exponential growth rate of volumes. In addition we establish the persistence of the stability of a linear impulsive differential equation under sufficiently small nonlinear perturbations.     

Sufficient conditions for the asymptotic stability of nonlinear multidelay differential equations with linear parts defined by pairwise permutable matrices

In this paper a generalization of the delayed exponential defined by Khusainov and Shuklin (2003) [1] for autonomous linear delay systems with one delay defined by permutable matrices is given for delay systems with multiple delays and pairwise permutable matrices. Using this multidelay-exponential a solution of a Cauchy initial value problem is represented. By an application of this representation and using Pinto’s integral inequality an asymptotic stability results for some classes of nonlinear multidelay differential equations are proved.     

Oscillations and limit cycles in Lotka-Volterra systems with delays

In this paper, competitive Lotka-Volterra systems are studied that have distributed delays and constant coefficients on interaction terms and have time dependent growth rate vectors with an asymptotically constant average. Algebraic conditions are found to rule out non-vanishing oscillations for such systems and heteroclinic limit cycles for autonomous systems. As a supplement to these results, simple sufficient conditions are provided for certain components of all solutions to vanish and a criterion is given for partial permanence. An outstanding feature of all these results is that the conditions are irrelevant of the size and distribution of the delays.