Global stability and bifurcation analysis of a delay induced prey-predator system with stage structure

This paper describes a delay induced prey–predator system with stage structure for prey. The dynamical characteristics of the system are rigorously studied using mathematical tools. The coexistence equilibria of the system is determined and the dynamic behavior of the system is investigated around coexistence equilibria. Sufficient conditions are derived for the global stability of the system. The optimal harvesting problem is formulated and solved in order to achieve the sustainability of the system, keeping the ecological balance, and maximize the monetary social benefit. Maturation time delay of prey is incorporated and the existence of Hopf bifurcation phenomenon is examined at the coexistence equilibria. It is shown that the time delay can cause a stable equilibrium to become unstable and even a switching of stabilities. Moreover, we use normal form method and center manifold theorem to examine the nature of the Hopf bifurcation. Finally, some numerical simulations are given to verify the analytical results, and the system is analyzed through graphical illustrations.     

Nonlinear dynamics of a delayed Leslie predator–prey model

The present paper is concerned with a delayed Leslie predator–prey model. The conditions of boundedness of the solutions of the system, existence, and stability of the equilibrium of the system are investigated. Meanwhile, we find that the system can also undergo a Hopf bifurcation of nonconstant periodic solution at the positive equilibrium when the delay crosses through a sequence of critical values. The extensive simulations carried out show that the bifurcations arise around the positive equilibrium.     

Rich dynamics in a non-local population model over three patches

We consider a system of nonlinear delay differential equations that describes the growth of the mature population of a species with age-structure living over three patches. We analyze existence of non-negative homogeneous equilibria and their stability and discuss possible Hopf bifurcation from these equilibria. More precisely, by employing both the standard Hopf bifurcation theory and the symmetric bifurcation theory for functional differential equations, we obtain very rich dynamics for the system, including bistable equilibria, transient oscillations, synchronous periodic solutions, phase-locked periodic solutions, mirror-reflecting waves and standing waves.     

Stability and Hopf bifurcation in a three-neuron unidirectional ring with distributed delays

In this paper, we consider the effect of distributed delays in a three-neuron unidirectional ring. Sufficient conditions for existence of unique equilibrium, multiple equilibria and their local stability are derived. Taking the average delay as a bifurcation parameter, we find two critical values at which the system undergoes Hopf bifurcations. The orbital asymptotic stability of the Hopf bifurcating periodic solutions is investigated by using the method of multiple scales. The global Hopf bifurcation is also studied. Finally, the theoretical results are illustrated by some numerical simulations.     

Bifurcation analysis of 4-axle rail vehicle models in a curved track

The dynamics of a diffusive predator–prey model with time delay and Michaelis–Menten-type harvesting subject to Neumann boundary condition is considered. Turing instability and Hopf bifurcation at positive equilibrium for the system without delay are investigated. Time delay-induced instability and Hopf bifurcation are also discussed. By the theory of normal form and center manifold, conditions for determining the bifurcation direction and the stability of bifurcating periodic solution are derived. Some numerical simulations are carried out for illustrating the theoretical results.     

Stability and bifurcation in a generalized delay prey–predator model

The present paper considers a generalized prey–predator model with time delay. It studies the stability of the nontrivial positive equilibrium and the existence of Hopf bifurcation for this system by choosing delay as a bifurcation parameter and analyzes the associated characteristic equation. The researcher investigates the direction of this bifurcation by using an explicit algorithm. Eventually, some numerical simulations are carried out to support the analytical results.     

Dynamics in two nonsmooth predator–prey models with threshold harvesting

Considering a good pest control program should reduce the pest to levels acceptable to the public, we investigate the threshold harvesting policy on pests in two predator–prey models. Both models are nonsmooth and the aim of this paper is to provide how threshold harvesting affects the dynamics of the two systems. When the harvesting threshold is larger than some positive level, the harvesting does not affect the ecosystem; when the harvesting threshold is less than the level, the model has complex dynamics with multiple coexistence equilibria, limit cycle, bistability, homoclinic orbit, saddle-node bifurcation, transcritical bifurcation, subcritical and supercritical Hopf bifurcation, Bogdanov–Takens bifurcation, and discontinuous Hopf bifurcation. Firstly, we provide the complete stability analysis and bifurcation analysis for the two models. Furthermore, some numerical simulations are given to illustrate our results. Finally, it is found that harvesting lowers the level of both species for natural enemy–pest system while raises the densities of both species for the pest–crop system. It is seen that the threshold harvesting policy of the enemy system is more effective than the crop system.     

Bifurcation and chaos of a delayed predator-prey model with dormancy of predators

In this paper, a delayed predator-prey model with dormancy of predators is investigated. It shows that time delay in the prey-species growth can lead to the occurrence of Hopf bifurcation with stability switches at a coexistence equilibrium. The computing formulas of stability and direction of the Hopf bifurcating periodic solutions are given. Under appropriate conditions, the uniform persistence of this model with time delay is proved. In this simple model, multiple periodic solutions coexist. Through numerical simulation, it is shown that different values of time delay can generate or eliminate chaos. Biologically, our results imply that dynamical behaviors of this system with time delay strongly depend on the initial density of this model and the time delay of the growth of the prey.     

Multiple scales and normal forms in a ring of delay coupled oscillators with application to chaotic Hindmarsh–Rose neurons

We study the appearance and stability of spatiotemporal periodic patterns like phase-locked oscillations, mirror-reflecting waves, standing waves, in-phase or antiphase oscillations, and coexistence of multiple patterns, in a ring of bidirectionally delay coupled oscillators. Hopf bifurcation, Hopf–Hopf bifurcation, and the equivariant Hopf bifurcation are studied in the viewpoint of normal forms obtained by using the method of multiple scales which is a kind of perturbation technique, thus a clear bifurcation scenario is depicted. We find time delay significantly affects the dynamics and induces rich spatiotemporal patterns. With the help of the unfolding system near Hopf–Hopf bifurcation, it is confirmed in some regions two kinds of stable oscillations may coexist. These phenomena are shown for the delay coupled limit cycle oscillators as well as for the delay coupled chaotic Hindmarsh–Rose neurons.     

A delayed ratio-dependent predator–prey model of interacting populations with Holling type III functional response

In this article, we study a ratio-dependent predator–prey model described by a Holling type III functional response with time delay incorporated into the resource limitation of the prey logistic equation. This investigation includes the influence of intra-species competition among the predator species. All the equilibria are characterized. Qualitative behavior of the complicated singular point (0,0) in the interior of the first quadrant is investigated by means of a blow-up transformation. Uniform persistence, stability, and Hopf bifurcation at the positive equilibrium point of the system are examined. Global asymptotic stability analyses of the positive equilibrium point by the Bendixon–Dulac criterion for non-delayed model and by constructing a suitable Lyapunov functional for the delayed model are carried out separately. We perform a numerical simulation to validate the applicability of the proposed mathematical model and our analytical findings.     

van der Pol型时滞系统的两参数余维一Hopf分岔及其稳定性

研究具有三次非线性时滞项的ｖａｎ ｄｅｒ Ｐｏｌ型时滞系统随两参数（时滞量和增益系数）余维一Ｈｏｐｆ分岔,说明了线性化特性方程随两参数变化时的根的分布和Ｈｏｐｆ分岔存在性;通过构造中心流形并且使用范式方法确定出Ｈｏｐｆ分岔的方向以及周期解的稳定性;分析了时滞量对所论系统发生Ｈｏｐｆ分岔的影响。     

Effort dynamics of a delay-induced prey–predator system with reserve

This paper describes a prey?Cpredator fishery system with prey dispersal in a two-patch environment, one of which is a free fishing zone and the other a protected zone. The proposed system reflects the dynamic interaction between the net economic revenue and the fishing effort used to harvest the population in presence of a suitable tax. Local as well as global stability of the system is analyzed. The optimal taxation policy is formulated and solved with the help of Pontryagin??s maximal principle. The objective of the paper is to achieve the sustainability of the fishery, keeping the ecological balance, and maximize the monetary social benefit. The dynamical behavior of the delay system is further analyzed through incorporating discrete type gestational delay of predators, and the existence of Hopf bifurcation phenomenon is checked at the interior equilibrium point. Moreover, we use normal form method and center manifold theorem to examine the nature of the Hopf bifurcation. Theoretical results are verified with the help of numerical examples and graphical illustrations.     

Stability and bifurcation analysis of delay coupled Van der Pol–Duffing oscillators

In this paper, we aim to investigate the dynamics of a system of Van der Pol–Duffing oscillators with delay coupling. First, taking the time delay as a bifurcation parameter, the stability of the equilibrium, and the existence of Hopf bifurcation are investigated. Then using the center manifold reduction technique and normal form theory, we give the direction of the Hopf bifurcation. And then by means of the symmetric bifurcation theory for delay differential equations and the representation theory of groups, we claim the bifurcation periodic solution induced by time delay is antiphase locked oscillation. Finally, at the end of the paper, numerical simulations are carried out to support our theoretical analysis.     

Dynamics analysis of a non-smooth Filippov pest-natural enemy system with time delay

In this paper, we propose a non-smooth Filippov system that describes the interaction of the pest and natural enemy with considering time delay, which represents the change in the growth rate of natural enemies before it is released to prey on pests. When the number of the pest is below the threshold, no control is applied; otherwise, control measures will be adopted. We discuss the stability of the equilibria and the existence of Hopf bifurcation. The results show that the Hopf bifurcation occurs when the time delay passes through some critical values. By applying the Filippov convex method, we obtain the dynamics of the sliding mode. The solutions of the system eventually tend toward the regular equilibrium, the pseudo-equilibrium or a standard periodic solution. Numerical simulations show that time delay plays an important role in local and global sliding bifurcations. We can obtain boundary focus bifurcations from boundary node bifurcations by varying time delay. Furthermore, touching, buckling and crossing bifurcations can be obtained frequently by increasing time delay. The results can provide some insights in pest control.

     

Bifurcation analysis of a diffusive ratio-dependent predator–prey model

In this paper, a ratio-dependent predator–prey model with diffusion is considered. The stability of the positive constant equilibrium, Turing instability, and the existence of Hopf and steady state bifurcations are studied. Necessary and sufficient conditions for the stability of the positive constant equilibrium are explicitly obtained. Spatially heterogeneous steady states with different spatial patterns are determined. By calculating the normal form on the center manifold, the formulas determining the direction and the stability of Hopf bifurcations are explicitly derived. For the steady state bifurcation, the normal form shows the possibility of pitchfork bifurcation and can be used to determine the stability of spatially inhomogeneous steady states. Some numerical simulations are carried out to illustrate and expand our theoretical results, in which, both spatially homogeneous and heterogeneous periodic solutions are observed. The numerical simulations also show the coexistence of two spatially inhomogeneous steady states, confirming the theoretical prediction.     

A delayed predator–prey model with strong Allee effect in prey population growth

In this paper, we consider a delayed predator-prey system with intraspecific competition among predator and a strong Allee effect in prey population growth. Using the delay as bifurcation parameter, we investigate the stability of coexisting equilibrium point and show that Hopf-bifurcation can occur when the discrete delay crosses some critical magnitude. The direction of the Hopf-bifurcating periodic solution and its stability are determined by applying the normal form method and the centre manifold theory. In addition, special attention is paid to the global continuation of local Hopf bifurcations. Using the global Hopf-bifurcation result of Wu ({Trans. Am. Math. Soc.} 350:4799?C4838, 1998) for functional differential equations, we establish the global existence of periodic solutions. Numerical simulations are carried out to validate the analytical findings.     

Hopf bifurcations in a predator-prey system of population allelopathy with a discrete delay and a distributed delay

A delayed Lotka?CVolterra predator-prey system of population allelopathy with discrete delay and distributed maturation delay for the predator population described by an integral with a strong delay kernel is considered. By linearizing the system at the positive equilibrium and analyzing the associated characteristic equation, the asymptotic stability of the positive equilibrium is investigated and Hopf bifurcations are demonstrated. Furthermore, the direction of Hopf bifurcation and the stability of the bifurcating periodic solutions are determined by the normal form theory and the center manifold theorem for functional differential equations. Finally, some numerical simulations are carried out for illustrating the theoretical results.     

Stability and Hopf bifurcation analysis for an energy resource system

This paper investigates the dynamical behaviors for a four-dimensional energy resource system with time delay, especially in terms of equilibria analyses and Hopf bifurcation analysis. By setting the time delay as a bifurcation parameter, it is shown that Hopf bifurcation would occur when the time delay exceeds a sequence of critical values. Furthermore, the stability and direction of the Hopf bifurcation are determined via the normal form theory and the center manifold reduction theorem. Numerical examples are given in the end of the paper to verify the theoretical results.     

Stability and global Hopf bifurcation in toxic phytoplankton–zooplankton model with delay and selective harvesting

Consider that some zooplankton can be harvested for food and some phytoplankton can liberate toxin; a toxin producing phytoplankton–zooplankton model with delay and selective harvesting is proposed and investigated. We discuss the stability of equilibria and perform the analysis of Hopf bifurcation. More precisely, the global asymptotical stability of equilibria is investigated by the Lyapunov method and Dulac theorem. In addition, the computing formulas of stability and direction of the Hopf bifurcating periodic solutions are also given. Furthermore, we prove that there exists at least one positive periodic solution as a time delay varies in some regions by using the global Hopf-bifurcation result of Wu (Trans. Am. Math. Soc. 350:4799–4838, 1998) for functional differential equations. Finally, the impact of harvesting is discussed along with numerical results to provide some support to the analytical findings.     

Some new insights into the Liu system

The present work is devoted to giving new insights into the Liu chaotic system. The local dynamical entities, such as the number of equilibria, the stability of hyperbolic equilibria, and the stability of the nonhyperbolic equilibrium obtained by using the center manifold theorem, the pitchfork bifurcation, the degenerate pitchfork bifurcation, and Hopf bifurcations, are all analyzed when the parameters are varied in the space of parameters. All the closed orbits of the system are also proven rigorously to be nonplanar but only to be curves in space. Moreover, the existence of singularly degenerate heteroclinic cycles for a suitable choice of the parameters is investigated.