The impact of Allee effect on a predator–prey system with Holling type II functional response

In this paper, the Allee effect is incorporated into a predator–prey model with Holling type II functional response. Compared with the predator–prey model without Allee effect, we find that the Allee effect of prey species increases the extinction risk of both predators and prey. When the handling time of predators is relatively short and the Allee effect of prey species becomes strong, both predators and prey may become extinct. Moreover, it is shown that the model with Allee effect undergoes the Hopf bifurcation and heteroclinic bifurcation. The Allee effect of prey species can lead to unstable periodical oscillation. It is also found that the positive equilibrium of the model could change from stable to unstable, and then to stable when the strength of Allee effect or the handling time of predators increases continuously from zero, that is, the model admits stability switches as a parameter changes. When the Allee effect of prey species becomes strong, longer handling time of predators may stabilize the coexistent steady state.     

Analysis of a predator–prey model with modified Holling–Tanner functional response and time delay

In paper, a predator–prey model with modified Holling–Tanner functional response and time delay is discussed. It is proved that the system is permanent under some appropriate conditions. The local stability of the equilibria is investigated. By constructing a suitable Lyapunov functional, sufficient conditions are derived for the global stability of the positive equilibrium of the model.     

Global Analysis of Predator–Prey System with Hawk and Dove Tactics

Functional response of the Holling type II is incorporated into a predator–prey model with predators using hawk‐dove tactics to consider combination effects of nonlinear functional response and individual tactics. By mathematical analysis, it is shown that the model undergoes a sequence of bifurcations including saddle‐node bifurcation, supercritical Hopf bifurcation and homoclinic bifurcation. New phenomena are found that include the bistable coexistence of prey and predators in the form of a stable limit cycle and a stable positive equilibrium, the bistable coexistence of prey and predators in a large stable limit cycle that encloses three positive equilibria and a stable positive equilibrium within the cycle, and the bistable coexistence of two stable limit cycles.     

A Predator�CPrey Model with a Holling Type I Functional Response Including a Predator Mutual Interference

The most widely used functional response in describing predator–prey relationships is the Holling type II functional response, where per capita predation is a smooth, increasing, and saturating function of prey density. Beddington and DeAngelis modified the Holling type II response to include interference of predators that increases with predator density. Here we introduce a predator-interference term into a Holling type I functional response. We explain the ecological rationale for the response and note that the phase plane configuration of the predator and prey isoclines differs greatly from that of the Beddington–DeAngelis response; for example, in having three possible interior equilibria rather than one. In fact, this new functional response seems to be quite unique. We used analytical and numerical methods to show that the resulting system shows a much richer dynamical behavior than the Beddington–DeAngelis response, or other typically used functional responses. For example, cyclic-fold, saddle-fold, homoclinic saddle connection, and multiple crossing bifurcations can all occur. We then use a smooth approximation to the Holling type I functional response with predator mutual interference to show that these dynamical properties do not result from the lack of smoothness, but rather from subtle differences in the functional responses.     

Periodic solutions in non autonomous predator prey system with delays

In this paper, we consider a biological model for two predators and one prey with periodic delays. By assuming that one predator consumes prey according to Holling II functional response while the other predator consumes prey according to the Beddington–DeAngelis functional response, based on the coincidence degree theory, the existence of positive periodic solutions for this model is obtained under suitable conditions.     

The effect of prey refuge and time delay on a diffusive predator‐prey system with hyperbolic mortality

In this article, we investigate the effect of prey refuge and time delay on a diffusive predator‐prey system with Holling II functional response and hyperbolic mortality subject to Neumann boundary condition. More precisely, we study Turing instability of positive equilibrium by using refuge as parameter, instability and Hopf bifurcation induced by time delay. In addition, by the theory of normal form and center manifold, we derive conditions for determining the bifurcation direction and the stability of the bifurcating periodic solution. © 2016 Wiley Periodicals, Inc. Complexity 21: 446–459, 2016     

Ecological complexity and feedback control in a prey–predator system with Holling type III functional response

This article deals with a bioeconomic model of prey–predator system with Holling type III functional response. The dynamical behavior of the system is extensively discussed. Continuous type gestational delay of predators is incorporated in the system to study delay induced instability. It is observed that the system undergoes singularity induced bifurcation at interior equilibrium point when net economic revenue of the system increases through zero. State feedback controller is designed to stabilize the system at positive economic profit. Time delay is considered as a bifurcation parameter to prove the occurrence of Hopf bifurcation phenomenon in the neighborhood of the coexisting equilibrium point. Finally, some numerical simulations are carried out to verify the analytical results and the system is analyzed through graphical illustrations. © 2015 Wiley Periodicals, Inc. Complexity 21: 346–360, 2016     

Stability and bifurcation analysis of a prey–predator model with age based predation

It is observed that in large animals only adult predators take part in direct predation while suckling feed on milk of adult predators and juveniles are dependent on the dead prey stock killed by the adult predators. Some parts of the dead prey population is consumed by adult predators and remaining parts are consumed by juveniles and the remaining portion decays naturally. In light of this, a mathematical model is proposed to study the stability and bifurcation behaviour of a prey–predator system with age based predation. All the feasible equilibria of the system are obtained and the conditions for the existence of the interior equilibrium are determined. The local stability analysis of all the feasible equilibria is carried out and the possibility of Hopf-bifurcation of the interior equilibrium is studied. Finally, numerical simulation is conducted to support the analytical results.     

Chaos and Hopf bifurcation analysis for a two species predator–prey system with prey refuge and diffusion

In this paper, a delayed predator–prey model with Holling type II functional response incorporating a constant prey refuge and diffusion is considered. By analyzing the characteristic equation of linearized system corresponding to the model, we study the local asymptotic stability of the positive equilibrium of the system. By choosing the time delay due to gestation as a bifurcation parameter, the existence of Hopf bifurcations at the positive equilibrium is established. By applying the normal form and the center manifold theory, an explicit algorithm to determine the direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions is derived. Further, an example is presented to illustrate our main results. Finally, recurring to the numerical method, the influences of impulsive perturbations on the dynamics of the system are also investigated.     

Global dynamics of two phytoplankton-zooplankton models with toxic substances effect

In this paper, we investigate phytoplankton-zooplankton models with toxic substances effect and two different kinds of predator functional responses. For Holling type II predator functional response, it is shown that the local stability of the positive equilibrium implies global stability if there exists a unique positive equilibrium. When there exist multiple positive equilibria, the local stability of the positive equilibrium with small phytoplankton population density implies that the model occurs bistable phenomenon. These results also hold for Holling type III predator functional response under certain conditions.     

Turing pattern formation in a three species model with generalist predator and cross-diffusion

In a natural ecosystem, specialist predators feed almost exclusively on one species of prey. But generalist predators feed on many types of species. Consequently, their dynamics is not coupled to the dynamics of a specific prey population. However, the defense of prey formed by congregating made the predator tend to move in the direction of lower concentration of prey species. This is described by cross-diffusion in a generalist predator–prey model. First, the positive equilibrium solution is globally asymptotically stable for the ODE system and for the reaction–diffusion system without cross-diffusion, respectively, hence it does not belong to the classical Turing instability scheme. But it becomes linearly unstable only when cross-diffusion also plays a role. This implies that cross–diffusion can lead to the occurrence and disappearance of the instability. Our results exhibit some interesting combining effects of cross-diffusion, predations and intra-species interactions. Furthermore, we consider the existence and non-existence results concerning non-constant positive steady states (patterns) of the system. We demonstrate that cross-diffusion can create non-constant positive steady-state solutions.     

A modified Leslie–Gower predator–prey model with prey infection

The disease effect on ecological systems is an important issue from mathematical and experimental point of view. In this paper, we formulate and analyze a predator–prey model for the susceptible population, infected population and their predator population with modified Leslie–Gower (or Holling–Tanner) functional response. Mathematical analysis of the model equations with regard to invariance of nonnegativity and boundedness of solutions, local and global stability of the biological feasible equilibria and permanence of the system are presented. When the rate of infection crosses a critical value, we determine that the strictly positive interior equilibrium undergoes Hopf bifurcation. From our numerical simulations, we observe that the predation rate also plays an important role on the dynamic behavior of our system.     

Global asymptotical stability and persistent property for a diffusive predator–prey system with modified Leslie–Gower functional response

A diffusive predator–prey system with modified Holling–Tanner functional response and no-flux boundary condition is considered in this work. A sufficient condition which ensures persistence of the system is obtained. Furthermore, sufficient conditions for the global asymptotical stability of the unique positive equilibrium of the system are derived by using a comparison method. It is shown that our result supplements and complements one of the main results of Shi et al. [H.B. Shi, W.T. Li, G. Lin, Positive steady states of a diffusive predator–prey system with modified Holling–Tanner functional response, Nonlinear Analysis: Real World Applications 11 (2010) 3711–3721].     

Bifurcation analysis of a diffusive predator–prey system with nonconstant death rate and Holling III functional response

In this paper, a diffusive predator–prey system with Holling III functional response and nonconstant death rate subject to Neumann boundary condition is considered. We study the stability of equilibria, and Turing instability of the positive equilibrium. We also perform a detailed Hopf bifurcation analysis to PDE system, and derive conditions for determining the bifurcation direction and the stability of the bifurcating periodic solution. In addition, some numerical simulations are carried out.     

Nonconstant Positive Steady States and Pattern Formation of 1D Prey-Taxis Systems

Prey-taxis is the process that predators move preferentially toward patches with highest density of prey. It is well known to have an important role in biological control and the maintenance of biodiversity. To model the coexistence and spatial distributions of predator and prey species, this paper concerns nonconstant positive steady states of a wide class of prey-taxis systems with general functional responses over 1D domain. Linearized stability of the positive equilibrium is analyzed to show that prey-taxis destabilizes prey–predator homogeneity when prey repulsion (e.g., due to volume-filling effect in predator species or group defense in prey species) is present, and prey-taxis stabilizes the homogeneity otherwise. Then, we investigate the existence and stability of nonconstant positive steady states to the system through rigorous bifurcation analysis. Moreover, we provide detailed and thorough calculations to determine properties such as pitchfork and turning direction of the local branches. Our stability results also provide a stable wave mode selection mechanism for thee reaction–advection–diffusion systems including prey-taxis models considered in this paper. Finally, we provide numerical studies of prey-taxis systems with Holling–Tanner kinetics to illustrate and support our theoretical findings. Our numerical simulations demonstrate that the \(2\times 2\) prey-taxis system is able to model the formation and evolution of various striking patterns, such as spikes, periodic oscillations, and coarsening even when the domain is one-dimensional. These dynamics can model the coexistence and spatial distributions of interacting prey and predator species. We also give some insights on how system parameters influence pattern formation in these models.     

Non-constant positive steady state of a diffusive Leslie-Gower type food web system

A three species food web comprising of two preys and one predator in an isolated homogeneous habitat is considered. The preys are assumed to grow logistically. The predator follows modified Leslie-Gower dynamics and feeds upon the prey species according to Holling Type II functional response. The local stability of the constant positive steady state of the corresponding temporal system and the spatio-temporal system are discussed. The existence and non-existence of non-constant positive steady states are investigated.     

Positive steady states of a diffusive predator–prey system with modified Holling–Tanner functional response

In this paper, we study a diffusive predator–prey system with modified Holling–Tanner functional response under homogeneous Neumann boundary condition. The qualitative properties, including the global attractor, persistence property, local and global asymptotic stability of the unique positive constant equilibrium are obtained. We also establish the existence and nonexistence of nonconstant positive steady states of this reaction–diffusion system, which indicates the effect of large diffusivity.     

A predator-prey system with L data

In this paper, we are concerned with a system of nonlinear partial differential equations modeling a predator-prey system in heterogeneous habitats. Preys are assumed to follow a logistic growth in the absence of predation, and predators are assumed to feed on preys with a Holling type II functional response to prey density. Also, interactions between predators are modelized by the statement of a food pyramid condition. Assuming no-flux boundary conditions and L¹ data, we prove the existence of at least one weak solution.     

Global dynamics of a predator–prey model with time delay and stage structure for the prey

A stage-structured predator–prey system with Holling type-II functional response and time delay due to the gestation of predator is investigated. By analyzing the characteristic equations, the local stability of each of feasible equilibria of the system is discussed and the existence of a Hopf bifurcation at the coexistence equilibrium is established. By means of the persistence theory on infinite dimensional systems, it is proven that the system is permanent if the coexistence equilibrium exists. By using Lyapunov functionals and LaSalle invariant principle, it is shown that the trivial equilibrium is globally stable when both the predator-extinction equilibrium and the coexistence equilibrium are not feasible, and that the predator-extinction equilibrium is globally asymptotically stable if the coexistence equilibrium does not exist, and sufficient conditions are derived for the global stability of the coexistence equilibrium. Numerical simulations are carried out to illustrate the main theoretical results.     

Global dynamics of a delayed eco‐epidemiological model with Holling type‐III functional response

In this paper, an eco‐epidemiological model with Holling type‐III functional response and a time delay representing the gestation period of the predators is investigated. In the model, it is assumed that the predator population suffers a transmissible disease. The disease basic reproduction number is obtained. By analyzing the corresponding characteristic equations, the local stability of each of feasible equilibria and the existence of Hopf bifurcations at the disease‐free equilibrium and the endemic‐coexistence equilibrium are established, respectively. By using the persistence theory on infinite dimensional systems, it is proved that if the disease basic reproduction number is greater than unity, the system is permanent. By means of Lyapunov functionals and LaSalle's invariance principle, sufficient conditions are obtained for the global stability of the endemic‐coexistence equilibrium, the disease‐free equilibrium and the predator‐extinction equilibrium of the system, respectively. Copyright © 2013 John Wiley & Sons, Ltd.