Optimal impulsive harvesting policy for single population

In this paper, we established the exploitation of impulsive harvesting single autonomous population model by Logistic equation. By some special methods, we analysis the impulsive harvesting population equation and obtain existence, the explicit expression and global attractiveness of impulsive periodic solutions for constant yield harvest and proportional harvest. Then, we choose the maximum sustainable yield as management objective, and investigate the optimal impulsive harvesting policies respectively. The optimal harvest effort that maximizes the sustainable yield per unit time, the corresponding optimal population levels are determined. At last, we point out that the continuous harvesting policy is superior to the impulsive harvesting policy, however, the latter is more beneficial in realistic operation.     

食饵具脉冲扰动与捕食者具连续收获的时滞捕食-食饵模型

本文讨论了与生物资源管理相关的食饵具脉冲扰动与捕食者具连续收获的时滞捕食-食饵模型,得到了捕食者灭绝周期解的全局吸引和系统持久的充分条件.也证明了系统的所有解的一致完全有界.我们的结论为现实的生物资源管理提供了可靠的策略依据,也丰富了脉冲时滞微分方程的理论.     

OPTIMAL IMPULSIVE CONTROL IN COMPETITIVE SYSTEM

In this paper,the impulsive exploitation of two species periodic competitive system is considered.First,we show that this type of system with impulsive har- vesting has a unique positive periodic solution,which is globally asymptotically stable.Further,by choosing the maximum total revenues as the management objective,we investigate the optimal harvesting policies for periodic competi- tive system with impulsive harvesting.Finally,we obtain the optimal time to harvest and optimal population level.     

ON THE IMPULSIVE BEVERTON–HOLT EQUATION AND EXTINCTION CONDITIONS IN POPULATION DYNAMICS

Abstract This paper is devoted to investigate extinction and nonextinction conditions of the extended Beverton–Holt equation (BHE) for dynamics of populations in Ecology when potential discontinuities at sampling points are included in the model. The proposed model is described by means of four sequences of parameters. Two of them are the intrinsic growth rate and the carrying capacity sequences which are included in the basic BHE model. The other two ones, namely, the harvesting (i.e., the hunting or fishing quota) and the internal consumption (which can include positive and negative migrations in the considered population habitat) sequences are included to parameterize the model discontinuities. Such discontinuities are related to impulses in the corresponding continuous‐time logistic equation. The obtained results establish how the harvesting quota and/or the internal consumption has to be fixed to guarantee the population nonextinction or, eventually, its extinction. Finally, some controllability results related to the search of a carrying capacity sequence such that the solution of the proposed impulsive BHE tracks a reference one are obtained.     

Dynamic analysis of a pest management SEI model with saturation incidence concerning impulsive control strategy

According to biological strategy for pest control, we investigate the dynamic behavior of a pest management SEI model with saturation incidence concerning impulsive control strategy-periodic releasing infected pests at fixed times. We prove that all solutions of the system are uniformly ultimately bounded and there exists a globally asymptotically stable pest-eradication periodic solution when the impulsive period is less than some critical value. When the impulsive period is larger than some critical value, the stability of the pest-eradication periodic solution is lost; the system is uniformly permanent. Thus, we can use the stability of the positive periodic solution and its period to control insect pests at acceptably low levels. Numerical results show that the system we consider can take on various kinds of periodic fluctuations and several types of attractor coexistence and is dominated by period-doubling cascade, symmetry-breaking pitchfork bifurcation, quasi-periodic oscillate, chaos, and non-unique dynamics.     

Qualitative analysis and applications of a kind of state-dependent impulsive differential equations

This paper analyzes a certain type of impulsive differential equations (IDEs). Several useful theorems for its periodic solutions and their stabilities are given. The key idea is that a periodically time-dependent IDE can be transformed into the state-dependent IDE. As applications of our theory, the optimization problems in population dynamics are studied. That is, the maximum sustainable yields of single population models with periodically impulsive constant harvesting are discussed. Furthermore, we apply these results to the studies of the order-1 periodic solutions and their stability of a single population model with stage structure in which the mature is impulsively proportionally harvested while the immature is impulsively added with the constant.     

Impulsive perturbations in a periodic delay differential equation model of plankton allelopathy

In this paper, we consider the dynamic behavior of an impulsive delay differential equation model with the effects of interspecific allelopathic interaction. A good understanding of the permanence, extinction and the existence of positive periodic solutions is gained. It turns out that the impulsive controls play a crucial role in shaping the above dynamics of the system. Numerical simulations are presented to substantiate the analytical results.     

Evolutionary dynamics in a Lotka–Volterra competition model with impulsive periodic disturbance

In this paper, we develop a theoretical framework to investigate the influence of impulsive periodic disturbance on the evolutionary dynamics of a continuous trait, such as body size, in a general Lotka–Volterra‐type competition model. The model is formulated as a system of impulsive differential equations. First, we derive analytically the fitness function of a mutant invading the resident populations when rare in both monomorphic and dimorphic populations. Second, we apply the fitness function to a specific system of asymmetric competition under size‐selective harvesting and investigate the conditions for evolutionarily stable strategy and evolutionary branching by means of critical function analysis. Finally, we perform long‐term simulation of evolutionary dynamics to demonstrate the emergence of high‐level polymorphism. Our analytical results show that large harvesting effort or small impulsive harvesting period inhibits branching, while large impulsive harvesting period promotes branching. Copyright © 2015 John Wiley & Sons, Ltd.     

周期Gompertz生态系统中的最优脉冲控制收获策略

以周期Gompertz系统为基础,讨论了周期变化的单种群生物资源的收获优化问题及种群的动力学性质.在单位收获努力量假设下,以最大可持续收获量为管理目标,确定了线性收获下的最优收获策略,获得了最优收获努力量、最大可持续收获及相应的最优种群水平的显示表达式,为自然资源的开发和利用提供了理论依据.     

Homoclinic bifurcation for a general state-dependent Kolmogorov type predator–prey model with harvesting

In this paper, a general Kolmogorov type predator–prey model is considered. Together with a constant-yield predator harvesting, the state dependent feedback control strategies which take into account the impulsive harvesting on predators as well as the impulsive stocking on the prey are incorporated in the process of population interactions. We firstly study the existence of an order-1 homoclinic cycle for the system. It is shown that an order-1 positive periodic solution bifurcates from the order-1 homoclinic cycle through a homoclinic bifurcation as the impulsive predator harvesting rate crosses some critical value. The uniqueness and stability of the order-1 positive periodic solution are derived by applying the geometry theory of differential equations and the method of successor function. Finally, some numerical examples are provided to illustrate the main results. These results indicate that careful management of resources and harvesting policies is required in the applied conservation and renewable resource contexts.     

Positive solutions of a nonlinear impulsive equation with periodic boundary conditions

In this paper, we investigate nonlinear second order differential equations subject to linear impulse conditions and periodic boundary conditions. Sign properties of an associated Green’s function are exploited to get existence results for positive solutions of the nonlinear boundary value problem with impulse. Upper and lower bounds for positive solutions are also given. The results obtained yield periodic positive solutions of the corresponding periodic impulsive nonlinear differential equation on the whole real axis.     

收获捕食者的时滞脉冲捕食模型

讨论了与可再生生物资源管理相关的食饵具脉冲扰动与捕食者具连续收获的阶段结构时滞捕食-食饵模型,得到了捕食者灭绝周期解的全局吸引和系统持久的充分条件.也证明了系统的所有解的一致完全有界.结论为现实的可再生生物资源管理提供了可靠的策略依据.     

An impulsive controlled eco-epidemic model with disease in the prey

In this paper, we investigate the dynamic behavior of an eco-epidemic model with impulsive control strategy. By using Floquet theorem of impulsive differential equation, we show there is a globally stable prey eradication periodic solution when the impulsive period is less than some critical value. We study the permanence of the system. Numerical simulations show that the complex dynamics of the system depends on the values of impulsive period and impulsive perturbation, for example double period, triple period solutions.     

Hybrid approach for pest control with impulsive releasing of natural enemies and chemical pesticides: A plant–pest–natural enemy model

The agricultural pests can be controlled effectively by simultaneous use (i.e., hybrid approach) of biological and chemical control methods. Also, many insect natural enemies have two major life stages, immature and mature. According to this biological background, in this paper, we propose a three tropic level plant–pest–natural enemy food chain model with stage structure in natural enemy. Moreover, impulsive releasing of natural enemies and harvesting of pests are also considered. We obtain that the system has two types of periodic solutions: plant–pest-extinction and pest-extinction using stroboscopic maps. The local stability for both periodic solutions is studied using the Floquet theory of the impulsive equation and small amplitude perturbation techniques. The sufficient conditions for the global attractivity of a pest-extinction periodic solution are determined by the comparison technique of impulsive differential equations. We analyze that the global attractivity of a pest-extinction periodic solution and permanence of the system are evidenced by a threshold limit of an impulsive period depending on pulse releasing and harvesting amounts. Finally, numerical simulations are given in support of validation of the theoretical findings.     

Dynamic behaviors of the periodic Lotka–Volterra competing system with impulsive perturbations

In this paper, we investigate a classical periodic Lotka–Volterra competing system with impulsive perturbations. The conditions for the linear stability of trivial periodic solution and semi-trivial periodic solutions are given by applying Floquet theory of linear periodic impulsive equation, and we also give the conditions for the global stability of these solutions as a consequence of some abstract monotone iterative schemes introduced in this paper, which will be also used to get some sufficient conditions for persistence. By using the method of coincidence degree, the conditions for the existence of at least one strictly positive (componentwise) periodic solution are derived. The theoretical results are confirmed by a specific example and numerical simulations. It shows that the dynamic behaviors of the system we consider are quite different from the corresponding system without pulses.     

Optimal impulsive harvesting on non-autonomous Beverton–Holt difference equations

Many recent advances in the theory of the optimal economic exploitation of renewable fish resources have been gained by applying optimal control theory. However, despite these successes, much less is known about how seasonal environments affect the maximum sustainable yield (MSY) (or population persistence) and any effects of relations between intensity and frequency of harvesting. Assuming that fish populations follow Beverton–Holt equations we investigated impulsive harvesting in seasonal environments, focusing on both economic aspects and resource sustainability. We first investigated the existence and stability of a periodic solution and its analytic formula, and then showed that the population persistence depends on the intensity and frequency of harvesting. With the MSY as a management objective, we investigated optimal impulsive harvesting policies. The optimal harvesting effort that maximizes the sustainable yield, the corresponding optimal population level, and the MSY are obtained by using discrete Euler–Lagrange equations and product formulae, and their explicit expressions were obtained in terms of the intrinsic growth rate, the carrying capacity, and the impulsive moments. These results imply that harvest timing is of crucial importance to the MSY. Since impulsive differential equations incorporate elements of continuous and discrete systems, we can apply all results obtained for Beverton–Holt equations with impulsive effects to periodic logistic equations with impulsive harvesting.     

A Holling II functional response food chain model with impulsive perturbations

In this paper, we investigate a three trophic level food chain system with Holling II functional responses and periodic constant impulsive perturbations of top predator. Conditions for extinction of predator as a pest are given. By using the Floquet theory of impulsive equation and small amplitude perturbation skills, we consider the local stability of predator eradication periodic solution. Further, influences of the impulsive perturbation on the inherent oscillation are studied numerically, which shows the rich dynamics (for example: period doubling, period halfing, chaos crisis) in the positive octant. The dynamics behavior is found to be very sensitive to the parameter values and initial value.     

Optimal impulsive harvesting on non-autonomous Beverton–Holt difference equations

Many recent advances in the theory of the optimal economic exploitation of renewable fish resources have been gained by applying optimal control theory. However, despite these successes, much less is known about how seasonal environments affect the maximum sustainable yield (MSY) (or population persistence) and any effects of relations between intensity and frequency of harvesting. Assuming that fish populations follow Beverton–Holt equations we investigated impulsive harvesting in seasonal environments, focusing on both economic aspects and resource sustainability. We first investigated the existence and stability of a periodic solution and its analytic formula, and then showed that the population persistence depends on the intensity and frequency of harvesting. With the MSY as a management objective, we investigated optimal impulsive harvesting policies. The optimal harvesting effort that maximizes the sustainable yield, the corresponding optimal population level, and the MSY are obtained by using discrete Euler–Lagrange equations and product formulae, and their explicit expressions were obtained in terms of the intrinsic growth rate, the carrying capacity, and the impulsive moments. These results imply that harvest timing is of crucial importance to the MSY. Since impulsive differential equations incorporate elements of continuous and discrete systems, we can apply all results obtained for Beverton–Holt equations with impulsive effects to periodic logistic equations with impulsive harvesting.     

一类具有收获率的脉冲Lotka-Volterra竞争合作系统的八个正概周期解

研究了一类具有收获率的脉冲Lotka-Volterra竞争合作系统的正概周期解.通过利用重合度理论延拓定理、概周期理论和不等式分析技巧,获得了系统至少存在8个正概周期解的充分条件,推广和改进了早期文献的相关结果.     

脉冲泛函微分方程的正周期解

运用Leray-Schauder不动点定理研究一类脉冲泛函微分方程的正周期解的存在性,获得了存在正周期解的充分条件,改进了已知文献中的结果.