Complex dynamics and switching transients in periodically forced Filippov prey–predator system

By employing threshold policy control (TPC) in combination with the definition of integrated pest management (IPM), a Filippov prey–predator model with periodic forcing has been proposed and studied, and the periodic forcing is affected by assuming a periodic variation in the intrinsic growth rate of the prey. This study aims to address how the periodic forcing and TPC affect the pest control. To do this, the sliding mode dynamics and sliding mode domain have been addressed firstly by using Utkin’s equivalent control method, and then the existence and stability of sliding periodic solution are investigated. Furthermore, the complex dynamics including multiple attractors coexistence, period adding sequences and chaotic solutions with respect to bifurcation parameters of forcing amplitude and economic threshold (ET) have been investigated numerically in more detail. Finally the switching transients associated with pest outbreaks and their biological implications have been discussed. Our results indicate that the sliding periodic solution could be globally stable, and consequently the prey or pest population can be controlled such that its density falls below the economic injury level (EIL). Moreover, the switching transients have both advantages and disadvantages concerning pest control, and the magnitude and frequency of switching transients depend on the initial values of both populations, forcing amplitude and ET.     

Permanence and extinction of periodic predator–prey systems in a patchy environment with delay

This paper studies two species predator–prey Lotka–Volterra type dispersal systems with periodic coefficients and infinite delays, in which the prey species can disperse among n-patches, but the predator species is confined to one patch and cannot disperse. Sufficient and necessary conditions of integrable form for the permanence, extinction and the existence of positive periodic solutions are established, respectively. Some well-known results on the nondelayed periodic predator–prey Lotka–Volterra type dispersal systems are improved and extended to the delayed case.     

On Nonautonomous Prey predator Patchy System

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Bifurcation analysis for a three-species predator-prey system with two delays

In this paper, a three-species predator-prey system with two delays is investigated. By choosing the sum τ of two delays as a bifurcation parameter, we first show that Hopf bifurcation at the positive equilibrium of the system can occur as τ crosses some critical values. Second, we obtain the formulae determining the direction of the Hopf bifurcations and the stability of the bifurcating periodic solutions by using the normal form theory and center manifold theorem. Finally, numerical simulations supporting our theoretical results are also included.     

Dynamics of a competitive Lotka-Volterra system with three delays

In this paper, a competitive Lotka-Volterra system with three delays is investigated. By choosing the sum τ of three delays as a bifurcation parameter, we show that in the above system, Hopf bifurcation at the positive equilibrium can occur as τ crosses some critical values. And we obtain the formulae determining direction of Hopf bifurcation and stability of the bifurcating periodic solutions by using the normal form theory and center manifold theorem. Finally, numerical simulations supporting our theoretical results are also included.     

Bifurcations for a predator-prey system with two delays

In this paper, a predator-prey system with two delays is investigated. By choosing the sum τ of two delays as a bifurcation parameter, we show that Hopf bifurcations can occur as τ crosses some critical values. By deriving the equation describing the flow on the center manifold, we can determine the direction of the Hopf bifurcations and the stability of the bifurcating periodic solutions. In addition, special attention is paid to the global continuation of local Hopf bifurcations. Using a global Hopf bifurcation result of [J. Wu, Symmetric functional differential equations and neural networks with memory, Trans. Amer. Math. Soc. 350 (1998) 4799-4838], we may show the global existence of periodic solutions.     

Dynamic behaviors of the periodic predator–prey system with distributed time delays and impulsive effect

In this paper, a periodic predator–prey system with distributed time delays and impulsive effect is investigated. By using the Floquet theory of linear periodic impulsive equation, some conditions for the linear stability of trivial periodic solution and semi-trivial periodic solutions are obtained. It is proved that the system can be permanent if all the trivial and semi-trivial periodic solutions are linearly unstable. We improve some results in Guo and Chen (2009) [1].     

Persistence and global stability of Bazykin predator–prey model with Beddington–DeAngelis response function

In this article, a predator–prey model of Beddington–DeAngelis type with discrete delay is proposed and analyzed. The essential mathematical features of the proposed model are investigated in terms of local, global analysis and bifurcation theory. By analyzing the associated characteristic equation, it is found that the Hopf bifurcation occurs when the delay parameter τ crosses some critical values. In this article, the classical Bazykin’s model is modified with Beddington–DeAngelis functional response. The parametric space under which the system enters into Hopf bifurcation for both delay and non-delay cases are investigated. Global stability results are obtained by constructing suitable Lyapunov functions for both the cases. We also derive the explicit formulae for determining the stability, direction and other properties of bifurcating periodic solutions by using normal form and central manifold theory. Our analytical findings are supported by numerical simulations. Biological implication of the analytical findings are discussed in the conclusion section.     

Persistence in nonautonomous predator–prey systems with infinite delays

This paper studies the general nonautonomous predator–prey Lotka–Volterra systems with infinite delays. The sufficient and necessary conditions of integrable form on the permanence and persistence of species are established. A very interesting and important property of two-species predator–prey systems is discovered, that is, the permanence of species and the existence of a persistent solution are each other equivalent. Particularly, for the periodic system with delays, applying these results, the sufficient and necessary conditions on the permanence and the existence of positive periodic solutions are obtained. Some well-known results on the nondelayed periodic predator–prey Lotka–Volterra systems are strongly improved and extended to the delayed case.     

A numerical study of the formation of spatial patterns in twospotted spider mites

The aim of this paper is to study the formation of spatial patterns in a predator–prey system with Tetranychus urticae as prey and Phytoseiulus persimilis as predator. Logistic Lotka–Volterra predator–prey equations are solved numerically with two different response functions, two initial conditions and one data set. The spatial patterns are generated by introducing diffusion-driven instability in the predator–prey system. Among all parameters involved in predator–prey equations, only the predator interference parameter is varied to generate diffusion-driven instability leading to spatial patterns of population density. Spatial patterns are further generated with the inclusion of prey-taxis in the predator–prey system. Routh–Hurwitz’s conditions for stability are used to create instability with prey-taxis in the system. It is shown that it is possible to generate spatial patterns with zero flux boundary conditions even in a smaller domain with a suitable value of the predator interference parameter or prey-taxis.     

Stability analysis of predator–prey system with migrating prey and disease infection in both species

We formulated and studied a predator–prey system with migrating prey and disease infection in both species. We used Lotka–Volterra type functional response. Mathematically, we analyzed the dynamics of the system such as existence of non negative equilibria, their stability. The basic reproduction number R₀ for the proposed mathematical model is calculated. Disease is endemic if R₀ > 1. Model is simulated by assuming hypothetical initial values and parameters.     

Uniqueness of periodic solutions of a nonautonomous density-dependent predator–prey system

In this present paper, we investigate the uniqueness of periodic solutions of a nonautonomous density-dependent and ratio-dependent predator–prey system, where not only the prey density dependence but also the predator density dependence are considered, such that the studied predator–prey system conforms to the realistically biological environment. We start with a sufficient condition for the permanence of the system and then construct a weaker sufficient condition by introducing a specific set, denoted as Γ. Based on this Γ and the Brouwer fixed-point theorem, we obtain the existence condition of positive periodic solutions. Moreover, since the uniqueness of positive periodic solutions can be ensured by global attractiveness, we alternatively introduce a sufficient condition for global attractiveness. Similarly, we also provide a sufficient condition for the uniqueness of non-negative periodic solutions.     

Spatially nonhomogeneous equilibrium in a reaction-diffusion system with distributed delay

We consider a reaction-diffusion system with general time-delayed growth rate and kernel functions. The existence and stability of the positive spatially nonhomogeneous steady-state solution are obtained. Moreover, taking minimal time delay τ as the bifurcation parameter, Hopf bifurcation near the steady-state solution is proved to occur at a critical value τ=τ₀. Especially, the Hopf bifurcation is forward and the bifurcated periodic solutions are stable on the center manifold. The general results are applied to competitive and cooperative systems with weak or strong kernel function respectively.     

一类具有阶段结构的非自治捕食-食饵系统的持久性与周期解

研究了一类具有阶段结构的非自治的捕食-食饵系统的渐近性质,得到了在适当条件下该系统的持久性,对应周期系统正周期解的存在性、唯一性以及全局渐近稳定性.     

Analysis of a Lotka–Volterra food chain chemostat with converting time delays

A model of the food chain chemostat involving predator, prey and growth-limiting nutrients is considered. The model incorporates two discrete time delays in order to describe the time involved in converting processes. The Lotka–Volterra type increasing functions are used to describe the species uptakes. In addition to showing that solutions with positive initial conditions are positive and bounded, we establish sufficient conditions for the (i) local stability and instability of the positive equilibrium and (ii) global stability of the non-negative equilibria. Numerical simulation suggests that the delays have both destabilizing and stabilizing effects, and the system can produce stable periodic solutions, quasi-periodic solutions and strange attractors.     

Complex dynamics of a Holling type II prey–predator system with state feedback control

The complex dynamics of a Holling type II prey–predator system with impulsive state feedback control is studied in both theoretical and numerical ways. The sufficient conditions for the existence and stability of semi-trivial and positive periodic solutions are obtained by using the Poincaré map and the analogue of the Poincaré criterion. The qualitative analysis shows that the positive periodic solution bifurcates from the semi-trivial solution through a fold bifurcation. The bifurcation diagrams, Lyapunov exponents, and phase portraits are illustrated by an example, in which the chaotic solutions appear via a cascade of period-doubling bifurcations. The superiority of the state feedback control strategy is also discussed.     

Dynamics of a Delayed Predator-prey System with Stage Structure for Predator and Prey

In this paper, a predator-prey system with two discrete delays and stage structure for both the predator and the prey is investigated. The dynamical behaviors such as local stability and local Hopf bifurcation are analyzed by regarding the possible combinations of the two delays as bifurcating parameter. Some explicit formulae determining the direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions are derived by using the normal form method and the center manifold theory. Finally, numerical simulations are presented to support the theoretical analysis.     

On the stability and Hopf bifurcation of a delay-induced predator–prey system with habitat complexity

We study the effect of the degree of habitat complexity and gestation delay on the stability of a predator–prey model. It is observed that there is stability switches, and Hopf bifurcation occurs when the delay crosses some critical value. By applying the normal form theory and the center manifold theorem, the explicit formulae which determine the stability and direction of the bifurcating periodic solutions are determined. The qualitative dynamical behavior of the model system is verified with the published data of Paramecium aurelia (prey) and Didinium nasutum (predator) interaction. It is observed that the quantitative level of abundance of system populations depends crucially on the delay parameter if the gestation period exceeds some critical value. However, the fluctuations in the population levels can be controlled completely by increasing the degree of habitat complexity.     

Periodic solutions of a discrete time nonautonomous ratio-dependent predator-prey system

With the help of differential equations with piecewise constant arguments, we first propose a discrete analogue of continuous time ratio-dependent predator-prey system, which is governed by nonautonomous difference equations, modeling the dynamics of the prey and the predator having nonoverlapping generations. Then, easily verifiable sufficient criteria are established for the existence of positive periodic solutions. The approach is based on the coincidence degree and the related continuation theorem as well as some priori estimates.