Bifurcation and control of a bioeconomic model of a prey–predator system with a time delay

In this paper, we analyze the dynamical behaviour of a bioeconomic model system using differential algebraic equations. The system describes a prey–predator fishery with prey dispersal in a two-patch environment, one of which is a free fishing zone and other is a protected zone. It is observed that a singularity-induced bifurcation phenomenon appears when a variation of the economic interest of harvesting is taken into account. We have incorporated a state feedback controller to stabilize the model system in the case of positive economic interest. A discrete-type gestational delay of predators is incorporated, and its effect on the dynamical behaviour of the model is analyzed. The occurrence of Hopf bifurcation of the proposed model with positive economic profit is shown in the neighbourhood of the coexisting equilibrium point through considering the delay as a bifurcation parameter. Finally, some numerical simulations are given to verify the analytical results, and the system is analyzed through graphical illustrations.     

Dynamical behavior in a stage-structured differential-algebraic prey-predator model with discrete time delay and harvesting

A differential-algebraic model system which considers a prey-predator system with stage structure for prey and harvest effort on predator is proposed. By using the differential-algebraic system theory and bifurcation theory, dynamic behavior of the proposed model system with and without discrete time delay is investigated. Local stability analysis of the model system without discrete time delay reveals that there is a phenomenon of singularity induced bifurcation due to variation of the economic interest of harvesting, and a state feedback controller is designed to stabilize the proposed model system at the interior equilibrium; Furthermore, local stability of the model system with discrete time delay is studied. It reveals that the discrete time delay has a destabilizing effect in the population dynamics, and a phenomenon of Hopf bifurcation occurs as the discrete time delay increases through a certain threshold. Finally, numerical simulations are carried out to show the consistency with theoretical analysis obtained in this paper.     

带有非线性收获和妊娠时滞的微分代数捕食者-食饵系统的稳定性及Hopf分支

本文研究了一类同时带有非线性食饵收获和捕食者妊娠时滞的微分代数捕食者-食饵系统的稳定性及Hopf分支问题.利用了分支理论和稳定性理论,以捕食者妊娠时滞作为系统的分支参数,获得了所提出的新系统在正平衡点处系统稳定性的相关判据条件和Hopf分支的产生条件.推广了一般带有线性收获和时滞的微分代数捕食者-食饵系统的结论.     

Delay-induced Bogdanov–Takens bifurcation in a Leslie–Gower predator–prey model with nonmonotonic functional response

This work is concerned with the dynamics of a Leslie–Gower predator–prey model with nonmonotonic functional response near the Bogdanov–Takens bifurcation point. By analyzing the characteristic equation associated with the nonhyperbolic equilibrium, the critical value of the delay inducing the Bogdanov–Takens bifurcation is obtained. In this case, the dynamics near this nonhyperbolic equilibrium can be reduced to the study of the dynamics of the corresponding normal form restricted to the associated two-dimensional center manifold. The bifurcation diagram near the Bogdanov–Takens bifurcation point is drawn according to the obtained normal form. We show that the change of delay can result in heteroclinic orbit, homoclinic orbit and unstable limit cycle.     

一类带无选择性捕获和时滞的捕食食饵系统的Hopf分支分析

本文主要研究了一个改进的带时滞和无选择捕获函数的捕食-食饵生态经济系统的稳定性和Hopf分支.利用微分代数系统的稳定性理论和分支理论,得到了系统正平衡点稳定性的条件,以及当时滞τ作为分支参数时系统产生Hopf分支的条件.对Leslie-Gower捕食-食饵模型进行了一定程度的完善,使得建立的模型更符合实际情况,因此得到的结论也更加科学.     

Hopf bifurcations in a predator–prey system with a discrete delay and a distributed delay

A delayed Lotka–Volterra two-species predator–prey system with discrete hunting delay and distributed maturation delay for the predator population described by an integral with a strong delay kernel is considered. By linearizing the system at the positive equilibrium and analyzing the associated characteristic equation, the asymptotic stability of the positive equilibrium is investigated and Hopf bifurcations are demonstrated. It is found that under suitable conditions on the parameters the positive equilibrium is asymptotically stable when the hunting delay is less than a certain critical value and unstable when the hunting delay is greater than this critical value. Meanwhile, according to the Hopf bifurcation theorem for functional differential equations (FDEs), we find that the system can also undergo a Hopf bifurcation of nonconstant periodic solution at the positive equilibrium when the hunting delay crosses through a sequence of critical values. In particular, by applying the normal form theory and the center manifold reduction for FDEs, an explicit algorithm determining the direction of Hopf bifurcations and the stability of bifurcating periodic solutions occurring through Hopf bifurcations is given. Finally, to verify our theoretical predictions, some numerical simulations are also included at the end of this paper.     

Hopf bifurcation and multiple periodic solutions in Lotka–Volterra systems with symmetries

The purpose of this paper is to study Hopf bifurcations in a delayed Lotka–Volterra system with dihedral symmetry. By treating the response delay as bifurcation parameter and employing equivariant degree method, we obtain the existence of multiple branches of nonconstant periodic solutions through a local Hopf bifurcation around an equilibrium. We find that competing coefficients and the response delay in the system can affect the spatio-temporal patterns of bifurcating periodic solutions. According to their symmetric properties, a topological classification is given for these periodic solutions. Furthermore, an estimation is presented on minimal number of bifurcating branches. These theoretical results are helpful to better understand the complex dynamics induced by response delays and symmetries in Lotka–Volterra systems.     

Hopf bifurcation in a delayed differential–algebraic biological economic system

In this paper, we consider a differential–algebraic biological economic system with time delay where the model with Holling type II functional response incorporates a constant prey refuge and prey harvesting. By considering time delay as bifurcation parameter, we analyze the stability and the Hopf bifurcation of the differential–algebraic biological economic system based on the new normal form approach of the differential–algebraic system and the normal form approach and the center manifold theory. Finally, numerical simulations illustrate the effectiveness of our results.     

Dynamical analysis of a delayed singular prey–predator economic model with stochastic fluctuations

This article studies a delayed singular prey–predator economic model with stochastic fluctuations, which is described by differential‐algebraic equations due to a economic theory. Local stability and Hopf bifurcation condition are described on the delayed singular prey–predator economic model within deterministic environment. It reveals the sensitivity of the model dynamics on gestation time delay. A phenomenon of Hopf bifurcation occurs as the gestation time delay increases through a certain threshold. Subsequently, a singular stochastic prey–predator economic model with time delay is obtained by introducing Gaussian white noise terms to the above deterministic model system. The fluctuation intensity of population and harvest effort are calculated by Fourier transforms method. Numerical simulations are carried out to substantiate these theory analysis. © 2013 Wiley Periodicals, Inc. Complexity 19: 23–29, 2014     

Stability and Hopf bifurcation in a diffusive logistic population model with nonlocal delay effect

A reaction-diffusion model with logistic type growth, nonlocal delay effect and Dirichlet boundary condition is considered, and combined effect of the time delay and nonlocal spatial dispersal provides a more realistic way of modeling the complex spatiotemporal behavior. The stability of the positive spatially nonhomogeneous positive equilibrium and associated Hopf bifurcation are investigated for the case of near equilibrium bifurcation point and the case of spatially homogeneous dispersal kernel.     

Numerical time-delayed feedback control in a neural network with delay

In this article, by a nonstandard finite-difference method we obtain the general time delayed feedback control numerical discrete scheme for a delayed neural network model. Firstly, the local stability of the equilibria point is discussed according to the Neimark–Sacker bifurcation theory. Then, from the point of view of control, for any step-size, a general time delayed feedback control numerical algorithm is introduced to delay the onset of the Neimark–Sacker bifurcation at a desired point by choosing appropriate control parameters. This controller can deal with the general system that the natural equilibrium cannot be given by analytic expression. Finally, numerical examples are provided to illustrate the theoretical results. The results show that the time delayed feedback numerical scheme is better than a polynomial function time delayed feedback method.     

Saddle-node-Hopf bifurcation in a modified Leslie–Gower predator-prey model with time-delay and prey harvesting

From the saddle-node-Hopf bifurcation point of view, this paper considers a modified Leslie–Gower predator-prey model with time delay and the Michaelis–Menten type prey harvesting. Firstly, we discuss the stability of the equilibria, obtain the critical conditions for the saddle-node-Hopf bifurcation, and give the completion bifurcation set by calculating the universal unfoldings near the saddle-node-Hopf bifurcation point by using the normal form theory and center manifold theorem. Then we derive the parameter conditions for the existence of monostable coexistence equilibrium and the parameter regions in which both the prey-extinction and the coexistence equilibrium (or coexistence periodic or quasi-periodic solutions) are simultaneously stabilized. We also investigate the heteroclinic bifurcation, and describe the phenomenon that the periodic behavior disappears as through the heteroclinic bifurcation. Finally, some numerical simulations are performed to support our analytic results.     

Hopf bifurcation and stability analysis on discrete-time Hopfield neural network with delay

In this paper, a discrete-time Hopfield neural network with delay is considered. We give some sufficient conditions ensuring the local stability of the equilibrium point for this model. By choosing the delay as a bifurcation parameter, we demonstrated that Neimark–Sacker bifurcation (or Hopf bifurcation for map) would occur when the delay exceeds a critical value. A formula for determining the direction bifurcation and stability of bifurcation periodic solutions is given by applying the normal form theory and the center manifold theorem. Some numerical simulations for justifying the theoretical results are also provided.     

Dynamics of a modified Leslie-Gower model with gestation effect and nonlinear harvesting

This study focuses on the dynamics of a modified Leslie-Gower predator-prey model where the intake rate of prey is by per capita predator according to Crowley-Martin functional response and prey is harvested through nonlinear harvesting strategy. Further the time-delay $(\tau)$ is imposed to utilize gestation period of predations. We investigate the permanence analysis of proposed system. The local stability of non-delayed model at all possible equilibrium points is studied. It is shown that the given model undergoes Hopf bifurcation around positive equilibrium point with respect to delay parameter $\tau$. Subsequently the stability of Hopf bifurcation and its direction are explored through normal and center manifold theories. The derived theoretical results are justified with the help of numerical simulations.     

Dynamics in a diffusive predator–prey system with a constant prey refuge and delay

In this paper, a diffusive predator–prey system with a constant prey refuge and time delay subject to Neumann boundary condition is considered. Local stability and Turing instability of the positive equilibrium are studied. The effect of time delay on the model is also obtained, including locally asymptotical stability and existence of Hopf bifurcation at the positive equilibrium. And the properties of Hopf bifurcation are determined by center manifold theorem and normal form theorem of partial functional differential equations. Some numerical simulations are carried out.     

Stability and Hopf bifurcation of a modified delay predator-prey model with stage structure

In this paper, a modified delay predator-prey model with stage structure is established, which involves the economic factor and internal competition of all the prey and predator populations. By the methods of normal form and characteristic equation, we obtain the stability of the positive equilibrium point and the sufficient condition of the existence of Hopf bifurcation. We analyze the influence of the time delay on the equation and show the occurrence of Hopf bifurcation periodic solution. The simulation gives a visual understanding for the existence and direction of Hopf bifurcation of the model.     

Dynamic Hopf bifurcations generated by nonlinear terms

We consider the so-called delayed loss of stability phenomenon for singularly perturbed systems of differential equations in case that the associated autonomous system with a scalar parameter undergoes the Hopf bifurcation at the zero equilibrium point. It is assumed that the linearization of the associated system is independent of the parameter and the next terms in the expansion of the right-hand parts at zero are positive homogeneous of order α>1. Simple formulas are presented to estimate the asymptotic delay for the delayed loss of stability phenomenon. More precisely, we suggest sufficient conditions which ensure that zeros of a simple function ψ defined by the positive homogeneous nonlinear terms are the Hopf bifurcation points of the associated system, the sign of ψ at other points determines stability of the zero equilibrium, and the asymptotic delay equals the distance between the bifurcation point and a zero of some primitive of ψ.     

Bifurcation analysis of a modified Holling-Tanner predator-prey model with time delay

In this paper, a modified Holling-Tanner predator-prey model with time delay is considered. By regarding the delay as the bifurcation parameter, the local asymptotic stability of the positive equilibrium is investigated. Meanwhile, we find that the system can also undergo a Hopf bifurcation of nonconstant periodic solution at the positive equilibrium when the delay crosses through a sequence of critical values. In particular, we study the direction of Hopf bifurcation and the stability of bifurcated periodic solutions, an explicit algorithm is given by applying the normal form theory and the center manifold reduction for functional differential equations. Finally, numerical simulations supporting the theoretical analysis are also included.     

Chaos and Hopf bifurcation analysis for a two species predator–prey system with prey refuge and diffusion

In this paper, a delayed predator–prey model with Holling type II functional response incorporating a constant prey refuge and diffusion is considered. By analyzing the characteristic equation of linearized system corresponding to the model, we study the local asymptotic stability of the positive equilibrium of the system. By choosing the time delay due to gestation as a bifurcation parameter, the existence of Hopf bifurcations at the positive equilibrium is established. By applying the normal form and the center manifold theory, an explicit algorithm to determine the direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions is derived. Further, an example is presented to illustrate our main results. Finally, recurring to the numerical method, the influences of impulsive perturbations on the dynamics of the system are also investigated.     

Novel dynamics of a predator–prey system with harvesting of the predator guided by its population

A predator–prey model was extended to include nonlinear harvesting of the predator guided by its population, such that harvesting is only implemented if the predator population exceeds an economic threshold. The proposed model is a nonsmooth dynamic system with switches between the original predator-prey model (free subsystem) and a model with nonlinear harvesting (harvesting subsystem). We initially examine the dynamics of both the free and the harvesting subsystems, and then we investigate the dynamics of the switching system using theories of nonsmooth systems. Theoretical results showed that the harvesting subsystem undergoes multiple bifurcations, including saddle-node, supercritical Hopf, Bogdanov–Takens and homoclinic bifurcations. The switching system not only retains all of the complex dynamics of the harvesting system but also exhibits much richer dynamics such as a sliding equilibrium, sliding cycle, boundary node (saddle point) bifurcation, boundary saddle-node bifurcation and buckling bifurcation. Both theoretical and numerical results showed that, by implementing predator population guided harvesting, the predator and prey population could coexist in more scenarios than those in which the predator may go extinct for the continuous harvesting regime. They could either stabilize at an equilibrium or oscillate periodically depending on the value of the economic threshold and the initial value of the system.