Steady-state solutions of a reaction–diffusion system arising from intraguild predation and internal storage

Intraguild predation is added to a mathematical model of competition between two species for a single nutrient with internal storage in the unstirred chemostat. At first, we established the sharp a priori estimates for nonnegative solutions of the system, which assure that all of nonnegative solutions belong to a special cone. The selection of this special cone enables us to apply the topological fixed point theorems in cones to establish the existence of positive solutions. Secondly, existence for positive steady state solutions of intraguild prey and intraguild predator is established in terms of the principal eigenvalues of associated nonlinear eigenvalue problems by means of the degree theory in the special cone. It turns out that positive steady state solutions exist when the associated principal eigenvalues are both negative or both positive.     

一类具有种内竞争率的竞争扩散模型稳态解的存在性和稳定性

利用Lyapunov-Schmidt约化方法和扰动理论,研究了一类带有不同种内竞争率的两物种竞争扩散模型.首先给出了半平凡稳态解的存在性和稳定性的充分条件,表明在一定条件下新物种可以入侵.在此基础上进一步证实了共存态的存在性和稳定性,研究表明在适当条件下相互竞争的两物种能够共存.所得结果改进和完善了已有的工作.     

A microbial continuous culture system with diffusion and diversified growth

A reaction-diffusion model is presented to describe the microbial continuous culture with diversified growth. The existence of nonnegative solutions and attractors for the system is obtained, the stability of steady states and the steady state bifurcation are studied under three growth conditions. In the case of no growth inhibition or only product inhibition, the system admits one positive constant steady state which is stable; in the case of growth inhibition only by substrate, the system can have two positive constant steady states, explicit conditions of the stability and the steady state bifurcation are also determined. In addition, numerical simulations are given to exhibit the theoretical results.     

非均匀Chemostat中非单食物链模型稳态正解的存在性

研究一类由反应扩散方程组描述的非均匀Chemostat中微生物之间既表现竞争关系又表现捕食被捕食关系的模型．用特征值理论确定了系统正稳态解存在的必要条件，用锥映射不动点指数方法给出了系统正稳态解存在的充分条件．     

Coexistence and bistability of a competition model with mixed dispersal strategy

In this paper, we investigate a Lotka–Volterra competition model with Danckwerts boundary conditions in a one-dimensional habitat where one species assumes pure random diffusion while another one undergoes mixed movement (both random and directed movements). We focus on the joint influence of advection rate, intrinsic growth rate and interspecific competition coefficient on the competition outcomes. It turns out that there exist some critical curves which separate the stable region of the semitrivial steady states from the unstable one. The locations of these curves determine whether coexistence or bistability occurs. More precisely, there are various tradeoffs between advection rate, intrinsic growth rate and interspecific competition coefficient that allow the transition of competition outcomes including competition exclusion, coexistence and bistability. We illustrate our results in various parameter spaces.     

A discrete-time model with non-monotonic functional response and strong Allee effect in prey

In this paper, we investigate the impact of strong Allee effect on the stability of a discrete-time predator–prey model with a non-monotonic functional response. The dynamics of discrete-time predator–prey models with strong Allee effect is studied earlier. But, the mathematical investigations of predator–prey dynamics in discrete-time set up with Holling type-IV functional response and strong Allee effect in prey are lacking. The proposed model supports the coexistence of two steady states, and the mathematical features of the model are analyzed based on local stability and bifurcation theory. By considering the Allee parameter as the bifurcation parameter, we provide sufficient conditions for the flip and the Neimark–Sacker bifurcations. We observe that Allee parameter plays a significant role in the dynamics of the system.     

A competition un-stirred chemostat model with virus in an aquatic system

A reaction–diffusion system of two bacteria species competing a single limiting nutrient with the consideration of virus infection is derived and analysed. Firstly, the well-posedness of the system, the existence of the trivial and semi-trivial steady states, and some prior estimations of the steady states are given. Secondly, a single species subsystem with virus is studied. The stability of the trivial and semi-trivial steady states and the uniform persistence of the subsystem are obtained. Further, taking the infective ability of virus as a bifurcation parameter, the global structure of the positive steady states and the effect of virus on the positive steady states are established via bifurcation theory and limiting arguments. It shows that the backward bifurcation may occur. Some sufficient conditions for the existence, uniqueness and stability of the positive steady state are also obtained. Finally, some sufficient conditions on the existence of the positive steady states for the full system are derived by using the fixed point index theory. Some results on persistence or extinction for the full system are also obtained.     

Coexistence and stability of a competition—diffusion system in population dynamics

The coexistence and stability of the population densities of two competing species in a bounded habitat are investigated in the present paper, where the effect of dispersion (transportation) is taken into consideration. The mathematical problem involves a coupled system of Lotka-Volterra-type reaction-diffusion equations together with some initial and boundary conditions, including the Dirichlet, Neumann and third type. Necessary and sufficient conditions for the coexistence and competitive exclusion are established and the effect of diffusion is explicitly given. For the stability problem, general criteria for the stability and instability of a steady-state solution are established and then applied to various situations depending on the relative magnitude among the physical parameters. Also given are necessary and sufficient conditions for the existence of multiple steady-state solutions and the stability or instability of each of these solutions. Special attention is given to the Neumann boundary condition with respect to which some threshold results for the coexistence and stability or instability of the four uniform steady states are characterized. It is shown in this situation that only one of the four constant steady states is asymptotically stable while the remaining three are unstable. The stability or instability of these states depends solely on the relative magnitude among the various rate constants and is independent of the diffusion coefficients.     

Long-term coexistence for a competitive system of spatially varying gradient reaction–diffusion equations

Spatial distribution of interacting chemical or biological species is usually described by a system of reaction–diffusion equations. In this work we consider a system of two reaction–diffusion equations with spatially varying diffusion coefficients which are different for different species and with forcing terms which are the gradient of a spatially varying potential. Such a system describes two competing biological species. We are interested in the possibility of long-term coexistence of the species in a bounded domain. Such long-term coexistence may be associated either with a periodic in time solution (usually associated with a Hopf bifurcation), or with time-independent solutions. We prove that no periodic solution exists for the system. We also consider some steady states (the time-independent solutions) and examine their stability and bifurcations.     

Dynamical behaviors of a classical Lotka–Volterra competition–diffusion–advection system

This paper deals with a two-species competition model in a homogeneous advective environment, where two species are subjected to a net loss of individuals at the downstream end. Under the assumption that the advection and diffusion rates of two species are proportional, we give a basic classification on the global dynamics by employing the theory of monotone dynamical system. It turns out that bistability does not happen, but coexistence and competitive exclusion may occur. Furthermore, we present a complete classification on the global dynamics in terms of the growth rates of two species. However, once the above assumption does not hold, bistability may occur. In detail, there exists a tradeoff between growth rates of two species such that competition outcomes can shift between three possible scenarios, including competitive exclusion, bistability and coexistence. These results show that growth competence is important to determine dynamical behaviors.     

A competition model for two resources in un-stirred chemostat

This paper studies a un-stirred chemostat with two species competing for two growth-limiting, non-reproducing resources. We determine the conditions for positive steady states of the two species, and then consider the global attractors of the model. In addition, we obtain the conditions under which the two populations uniformly strongly persist or go to extinction. Since the diffusion mechanism with homogeneous boundary conditions inhibits the growth of the organism species, it can be understood that the coexistence will be ensured by proportionally smaller diffusions for the two species. In particular, it is found that both instability and bi-stability subcases of the two semitrivial steady states are included in the coexistence region. The two populations will go to extinction when both possess large diffusion rates. If just one of them spreads faster with the other one diffusing slower, then the related semitrivial steady state will be globally attracting. The techniques used for the above results consist of the degree theory, the semigroup theory, and the maximum principle.     

Bifurcation and stability of a two-species reaction–diffusion–advection competition model

This paper is concerned with the dynamics of a two-species reaction–diffusion–advection competition model subject to the no-flux boundary condition in a bounded domain. By the signs of the associated principal eigenvalues, we derive the existence and local stability of the trivial and semi-trivial steady-state solutions. Moreover, the nonexistence and existence of the coexistence steady-state solutions stemming from the two boundary steady states are obtained as well. In particular, we describe the feature of the coincidence of bifurcating coexistence steady-state solution branches. At the same time, the effect of advection on the stability of the bifurcating solution is also investigated, and our results suggest that the advection term may change the stability. Finally, we point out that the methods we applied here are mainly based on spectral analysis, perturbation theory, comparison principle, monotone theory, Lyapunov–Schmidt reduction, and bifurcation theory.     

Effects of prey over–undercrowding in predator–prey systems with prey-dependent trophic functions

The local dynamics of a two-trophic chain in the presence of both overcrowding and undercrowding effects on prey growth is investigated. The starting point is given by a general predator–prey system, in which the prey growth rate and the trophic interaction function are defined only by some properties determining their shapes; in particular, the prey growth function is assumed to model a strong Allee effect. A stability analysis of the system using the predation efficiency as bifurcation parameter is performed; conditions for the existence and stability of extinction and coexistence equilibrium states are determined, and peculiar features of the dynamics exhibited by the system are presented, with particular attention to limit cycles and bistability situations. Results are compared with those obtained when overcrowding and undercrowding effects are considered separately.     

Nonconstant Positive Steady States and Pattern Formation of 1D Prey-Taxis Systems

Prey-taxis is the process that predators move preferentially toward patches with highest density of prey. It is well known to have an important role in biological control and the maintenance of biodiversity. To model the coexistence and spatial distributions of predator and prey species, this paper concerns nonconstant positive steady states of a wide class of prey-taxis systems with general functional responses over 1D domain. Linearized stability of the positive equilibrium is analyzed to show that prey-taxis destabilizes prey–predator homogeneity when prey repulsion (e.g., due to volume-filling effect in predator species or group defense in prey species) is present, and prey-taxis stabilizes the homogeneity otherwise. Then, we investigate the existence and stability of nonconstant positive steady states to the system through rigorous bifurcation analysis. Moreover, we provide detailed and thorough calculations to determine properties such as pitchfork and turning direction of the local branches. Our stability results also provide a stable wave mode selection mechanism for thee reaction–advection–diffusion systems including prey-taxis models considered in this paper. Finally, we provide numerical studies of prey-taxis systems with Holling–Tanner kinetics to illustrate and support our theoretical findings. Our numerical simulations demonstrate that the \(2\times 2\) prey-taxis system is able to model the formation and evolution of various striking patterns, such as spikes, periodic oscillations, and coarsening even when the domain is one-dimensional. These dynamics can model the coexistence and spatial distributions of interacting prey and predator species. We also give some insights on how system parameters influence pattern formation in these models.     

Stability results for a nonlinear two-species competition model with size-structure

We formulate a system of integro-differential equations to model the dynamics of competition in a two-species community, in which the mortality, fertility and growth are sizedependent. Existence and uniqueness of nonnegative solutions to the system are analyzed. The existence of the stationary size distributions is discussed, and the linear stability is investigated by means of the semigroup theory of operators and the characteristic equation technique. Some sufficient conditions for asymptotical stability/instability of steady states are obtained. The resulting conclusion extends some existing results involving age-independent and age-dependent population models.     

On a Lotka-Volterra Competition Diffusion Model with Advection

In this paper, the author focuses on the joint effects of diffusion and advection on the dynamics of a classical two species Lotka-Volterra competition-diffusion-advection system, where the ratio of diffusion and advection rates are supposed to be a positive constant. For comparison purposes, the two species are assumed to have identical competition abilities throughout this paper. The results explore the condition on the diffusion and advection rates for the stability of former species. Meanwhile, an asymptotic behavior of the stable coexistence steady states is obtained.     

Competitive exclusion and coexistence for competitive systems on ordered Banach spaces

The dynamics of competitive maps and semiflows defined on the product of two cones in respective Banach spaces is studied. It is shown that exactly one of three outcomes is possible for two viable competitors. Either one or the other population becomes extinct while the surviving population approaches a steady state, or there exists a positive steady state representing the coexistence of both populations.

     

The effect of diffusion for the multispecies Lotka–Volterra competition model

The purpose of this paper is to study the effect of diffusion in the existence of non-constant steady states for the Lotka–Volterra competition-diffusion system with three species, under Neumann boundary conditions. It will be shown that two large diffusion rates prevent the appearance of non-constant steady states, while if just one species diffuses fast non-constant equilibria may arise. The existence is shown by two methods, degree theory and bifurcation techniques. The stability of bifurcating steady states will be established.     

Dynamics of a spatiotemporal model on populations in a polluted river

In this paper, we investigate the dynamics for a reaction–diffusion–advection system which models populations in a polluted river. More precisely, we study the stability of steady states, which yields sufficient conditions that lead to population persistence or extinction. Furthermore, some dependence of the stability of the toxicant-only steady state and the population-toxicant coexistence steady state on the model parameters are given.     

Uniqueness and stability of positive steady state solutions for a ratio-dependent predator–prey system with a crowding term in the prey equation

This paper deals with a ratio-dependent predator–prey system with a crowding term in the prey equation, where it is assumed that the coefficient of the functional response is less than the coefficient of the intrinsic growth rates of the prey species. We demonstrate some special behaviors of solutions to the system which the coexistence states of two species can be obtained when the crowding region in the prey equation only is designed suitably. Furthermore, we demonstrate that under some conditions, the positive steady state solution of the predator–prey system with a crowding term in the prey equation is unique and stable. Our result is different from those ones of the predator–prey systems without the crowding terms.