Dynamics of a delayed predator–prey model with predator migration

Based on the availability of prey and a simple predator–prey model, we propose a delayed predator–prey model with predator migration to describe biological control. We first study the existence and stability of equilibria. It turns out that backward bifurcation occurs with the migration rate as bifurcation parameter. The stability of the trivial equilibrium and the boundary equilibrium is delay-independent. However, the stability of the positive equilibrium may be delay-dependent. Moreover, delay can switch the stability of the positive equilibrium. When the positive equilibrium loses stability, Hopf bifurcation can occur. The direction and stability of Hopf bifurcation is derived by applying the center manifold method and the normal form theory. The main theoretical results are illustrated with numerical simulations.     

Dynamics analysis of a delayed viral infection model with logistic growth and immune impairment

In this paper, the dynamics behavior of a delayed viral infection model with logistic growth and immune impairment is studied. It is shown that there exist three equilibria. By analyzing the characteristic equations, the local stability of the infection-free equilibrium and the immune-exhausted equilibrium of the model are established. By using suitable Lyapunov functional and LaSalle invariant principle, it is proved that the two equilibria are globally asymptotically stable. In the following, the stability of the positive equilibrium is investigated. Furthermore, we investigate the existence of Hopf bifurcation by using a delay as a bifurcation parameter. Finally, numerical simulations are carried out to explain the mathematical conclusions.     

Stability and Hopf bifurcation analysis of an eco-epidemic model with a stage structure

In this paper, an eco-epidemiological model with a stage structure is considered. The asymptotical stability of the five equilibria, the existence of stability switches about positive equilibrium, is investigated. It is found that Hopf bifurcation occurs when the delay τ passes though a critical value. Some explicit formulae determining the stability and the direction of the Hopf bifurcation periodic solutions bifurcating from Hopf bifurcations are obtained by using the normal form theory and center manifold theory. Some numerical simulations for justifying the theoretical analysis are also provided. Finally, biological explanations and main conclusions are given.     

Modeling and analysis of a prey–predator system with disease in the prey

A three dimensional ecoepidemiological model consisting of susceptible prey, infected prey and predator is proposed and analysed in the present work. The parameter delay is introduced in the model system for considering the time taken by a susceptible prey to become infected. Mathematically we analyze the dynamics of the system such as, boundedness of the solutions, existence of non-negative equilibria, local and global stability of interior equilibrium point. Next we choose delay as a bifurcation parameter to examine the existence of the Hopf bifurcation of the system around its interior equilibrium. Moreover we use the normal form method and center manifold theorem to investigate the direction of the Hopf bifurcation and stability of the bifurcating limit cycle. Some numerical simulations are carried out to support the analytical results.     

Dynamical behavior of a delay virus dynamics model with CTL immune response

In this paper, the dynamical behavior of a virus dynamics model with CTL immune response and time delay is studied. Time delay is used to describe the time between the infected cell and the emission of viral particles on a cellular level. The effect of time delay on stability of the equilibria of the CTL immune response model has been studied and sufficient criteria for local asymptotic stability of the disease-free equilibrium, immune-free equilibrium and endemic equilibrium and global asymptotic stability of the disease-free equilibrium are given. Some conditions for Hopf bifurcation around immune-free equilibrium and endemic equilibrium to occur are also obtained by using the time delay as a bifurcation parameter. Numerical simulation with some hypothetical sets of data has been done to support the analytical findings.     

Stability and Hopf bifurcation in a predator–prey model with stage structure for the predator

A predator–prey system with stage structure for the predator and time delay due to the gestation of the predator is investigated. By analyzing the corresponding characteristic equations, the local stability of a positive equilibrium and two boundary equilibria of the system is discussed, respectively. Further, the existence of a Hopf bifurcation at the positive equilibrium is also studied. By using an iteration technique and comparison argument, respectively, sufficient conditions are derived for the global stability of the positive equilibrium and one of the boundary equilibria of the proposed system. As a result, the threshold is obtained for the permanence and extinction of the system. Numerical simulations are carried out to illustrate the main results.     

Bifurcation analysis of a modified Holling-Tanner predator-prey model with time delay

In this paper, a modified Holling-Tanner predator-prey model with time delay is considered. By regarding the delay as the bifurcation parameter, the local asymptotic stability of the positive equilibrium is investigated. Meanwhile, we find that the system can also undergo a Hopf bifurcation of nonconstant periodic solution at the positive equilibrium when the delay crosses through a sequence of critical values. In particular, we study the direction of Hopf bifurcation and the stability of bifurcated periodic solutions, an explicit algorithm is given by applying the normal form theory and the center manifold reduction for functional differential equations. Finally, numerical simulations supporting the theoretical analysis are also included.     

Stability and Hopf bifurcation of a delayed viral infection model with logistic growth and saturated immune impairment

In this paper, the stability and Hopf bifurcation of a delayed viral infection model with logistic growth and saturated immune impairment is studied. It is shown that there exist 3 equilibria. The sufficient conditions for local asymptotic stability of the infection‐free equilibrium and no‐immune equilibrium are given. We also discussed the local stability of positive equilibrium and the existence of Hopf bifurcation. Moreover, the direction and stability of Hopf bifurcation is obtained by using standard form theory and the center manifold theorem. Finally, numerical simulations are performed to verify the theoretical conclusions.     

一类具有阶段结构和时滞的捕食模型的稳定性及Hopf分支(英文)

研究一类具有阶段结构和时滞的捕食模型.通过特征方程分别分析了正平衡点和边界平衡点的局部稳定性,到了系统Hopf分支存在的充分条件.通过规范型理论和中心流型定理,给出了确定Hopf分支方向和分支周期解的稳定性的计算公式.     

Bifurcation analysis and chaos control of the modified Chua’s circuit system

From the view of bifurcation and chaos control, the dynamics of modified Chua’s circuit system are investigated by a delayed feedback method. Firstly, the local stability of the equilibria is discussed by analyzing the distribution of the roots of associated characteristic equation. The regions of linear stability of equilibria are given. It is found that there exist Hopf bifurcation and Hopf-zero bifurcation when the delay passes though a sequence of critical values. By using the normal form method and the center manifold theory, we derive the explicit formulas for determining the direction and stability of Hopf bifurcation. Finally, chaotic oscillation is converted into a stable equilibrium or a stable periodic orbit by designing appropriate feedback strength and delay. Some numerical simulations are carried out to support the analytic results.     

Bifurcation of an eco-epidemiological model with a nonlinear incidence rate

A predator-prey system with disease in the prey is considered. Assume that the incidence rate is nonlinear, we analyse the boundedness of solutions and local stability of equilibria, by using bifurcation methods and techniques, we study Bogdanov-Takens bifurcation near a boundary equilibrium, and obtain a saddle-node bifurcation curve, a Hopf bifurcation curve and a homoclinic bifurcation curve. The Hopf bifurcation and generalized Hopf bifurcation near the positive equilibrium is analyzed, one or two limit cycles is also discussed.     

Stability analysis in a diffusional immunosuppressive infection model with delayed antiviral immune response

In this paper, the diffusion is introduced to an immunosuppressive infection model with delayed antiviral immune response. The direction and stability of Hopf bifurcation are effected by time delay, in the absence of which the positive equilibrium is locally asymptotically stable by means of analyzing eigenvalue spectrum; however, when the time delay increases beyond a threshold, the positive equilibrium loses its stability via the Hopf bifurcation. The stability and direction of the Hopf bifurcation is investigated with the norm form and the center manifold theory. The stability of the Hopf bifurcation leads to the emergence of spatial spiral patterns. Numerical calculations are performed to illustrate our theoretical results. Copyright © 2017 John Wiley & Sons, Ltd.     

Hopf bifurcations in a predator–prey system with a discrete delay and a distributed delay

A delayed Lotka–Volterra two-species predator–prey system with discrete hunting delay and distributed maturation delay for the predator population described by an integral with a strong delay kernel is considered. By linearizing the system at the positive equilibrium and analyzing the associated characteristic equation, the asymptotic stability of the positive equilibrium is investigated and Hopf bifurcations are demonstrated. It is found that under suitable conditions on the parameters the positive equilibrium is asymptotically stable when the hunting delay is less than a certain critical value and unstable when the hunting delay is greater than this critical value. Meanwhile, according to the Hopf bifurcation theorem for functional differential equations (FDEs), we find that the system can also undergo a Hopf bifurcation of nonconstant periodic solution at the positive equilibrium when the hunting delay crosses through a sequence of critical values. In particular, by applying the normal form theory and the center manifold reduction for FDEs, an explicit algorithm determining the direction of Hopf bifurcations and the stability of bifurcating periodic solutions occurring through Hopf bifurcations is given. Finally, to verify our theoretical predictions, some numerical simulations are also included at the end of this paper.     

Stability and Hopf bifurcation of a delayed predator-prey system with nonlocal competition and herd behaviour

In this paper, we investigate the stability and Hopf bifurcation of a diffusive predator-prey system with herd behaviour. The model is described by introducing both time delay and nonlocal prey intraspecific competition. Compared to the model without time delay, or without nonlocal competition, thanks to the together action of time delay and nonlocal competition, we prove that the first critical value of Hopf bifurcation may be homogenous or non-homogeneous. We also show that a double-Hopf bifurcation occurs at the intersection point of the homogenous and non-homogeneous Hopf bifurcation curves. Furthermore, by the computation of normal forms for the system near equilibria, we investigate the stability and direction of Hopf bifurcation. Numerical simulations also show that the spatially homogeneous and non-homogeneous periodic patters.     

Delay induced subcritical Hopf bifurcation in a diffusive predator-prey model with herd behavior and hyperbolic mortality

In this paper, we consider the dynamics of a delayed diffusive predator-prey model with herd behavior and hyperbolic mortality under Neumann boundary conditions. Firstly, by analyzing the characteristic equations in detail and taking the delay as a bifurcation parameter, the stability of the positive equilibria and the existence of Hopf bifurcations induced by delay are investigated. Then, applying the normal form theory and the center manifold argument for partial functional differential equations, the formula determining the properties of the Hopf bifurcation are obtained. Finally, some numerical simulations are also carried out and we obtain the unstable spatial periodic solutions, which are induced by the subcritical Hopf bifurcation.     

Delay-induced oscillatory dynamics in humoral mediated immune response with two time delays

A mathematical model for the quantitative analysis of cancer immune interaction, considering the role of humoral (antibody) mediated immune response with two time delays, namely maturation and interaction delays has been proposed in this paper. The aim of this work is to assess the effect of time delays on the interaction between cancerous cells and the antibodies. After categorizing the parametric plane into different regions based on the existence of equilibria, we investigate both analytically and through simulations, the stability of equilibria and the onset of sustained oscillations through Hopf bifurcations. The direction and stability of the Hopf bifurcation which occurs at the positive interior equilibrium point of the system have also been studied. It is observed that both the delays play an important role in stability switching. Appropriate therapy with a proper choice of system parameters are suggested to obtain cancer free equilibrium.     

Stability and bifurcation analysis of a nutrient-phytoplankton model with time delay

We proposed a nutrient-phytoplankton interaction model with a discrete and distributed time delay to provide a better understanding of phytoplankton growth dynamics and nutrient-phytoplankton oscillations induced by delay. Standard linear analysis indicated that delay can induce instability of a positive equilibrium via Hopf bifurcation. We derived the conditions guaranteeing the existence of Hopf bifurcation and tracked its direction and the stability of the bifurcating periodic solutions. We also obtained the sufficient conditions for the global asymptotic stability of the unique positive steady state. Numerical analysis in the fully nonlinear regime showed that the stability of the positive equilibrium is sensitive to changes in delay values under select conditions. Numerical results were consistent with results predicted by linear analysis.     

Dynamics of a ratio‐dependent stage‐structured predator‐prey model with delay

In this paper, we investigate the dynamics of a time‐delay ratio‐dependent predator‐prey model with stage structure for the predator. This predator‐prey system conforms to the realistically biological environment. The existence and stability of the positive equilibrium are thoroughly analyzed, and the sufficient and necessary conditions for the stability and instability of the positive equilibrium are obtained for the case without delay. Then, the influence of delay on the dynamics of the system is investigated using the geometric criterion developed by Beretta and Kuang. 26 We show that the positive steady state can be destabilized through a Hopf bifurcation and there exist stability switches under some conditions. The formulas determining the direction and the stability of Hopf bifurcations are explicitly derived by using the center manifold reduction and normal form theory. Finally, some numerical simulations are performed to illustrate and expand our theoretical results.     

Dynamics in a diffusive predator–prey system with a constant prey refuge and delay

In this paper, a diffusive predator–prey system with a constant prey refuge and time delay subject to Neumann boundary condition is considered. Local stability and Turing instability of the positive equilibrium are studied. The effect of time delay on the model is also obtained, including locally asymptotical stability and existence of Hopf bifurcation at the positive equilibrium. And the properties of Hopf bifurcation are determined by center manifold theorem and normal form theorem of partial functional differential equations. Some numerical simulations are carried out.