Effect of prey-taxis on predator’s invasion in a spatially heterogeneous environment

In this study, we consider a diffusive predator–prey system with prey-taxis and ratio-dependent functional responses in a spatially heterogeneous environment. Prey-taxis implies that the predator exhibits directed movement in the presence of prey. We claim that when the predator diffuses uniformly without prey-taxis, only the diffusion of the species plays a role in the predator’s invasion in a spatially homogeneous environment. However, when the predator disperses through uniform diffusion together with prey-taxis, we observe that both the diffusion of species and prey-taxis affect the invasion of the predator, which is not the case in a spatially homogeneous environment. The results are obtained by investigating the local stability of a semi-trivial solution, using an eigenvalue analysis.     

Spatiotemporal complexity of a discrete space-time predator−prey system with self- and cross-diffusion

In this research, we investigate the spatiotemporal dynamics of a discrete space-time predator−prey system with self- and cross-diffusion. Through stability analysis and bifurcation analysis, Turing pattern formation conditions are derived and two nonlinear mechanisms of pattern formation are found, i.e., pure Turing instability and Hopf-Turing instability. Numerical simulations reveal rich dynamics of the discrete predator−prey system. In spatially homogeneous case, stable homogeneous stationary states, homogeneous periodic, quasiperiodic and chaotic oscillating states are exhibited; in spatially heterogeneous case, a surprising variety of prey and predator patterns are described, including spotted, striped, labyrinth, gapped, spiral, circled patterns and many intermediate patterns. Moreover, sensitivity of spatiotemporal pattern formation to initial conditions is predicted along with Hopf-Turing instability, suggesting the self-organization of diverse patterns under identical parametric conditions. In comparison with former results in literature, the discrete version of reaction-diffusion model developed in this research capture more complicated and richer nonlinear dynamical behaviors, contributing to a new comprehending on the complex pattern formation of spatially extended discrete predator−prey systems.     

Stability and Hopf bifurcation of a diffusive predator‐prey model with hyperbolic mortality

The dynamics of a reaction‐diffusion predator‐prey model with hyperbolic mortality and Holling type II response effect is considered. The stability of the positive equilibrium and the existence of Hopf bifurcation are investigated by analyzing the distribution of eigenvalues without diffusion. We also study the spatially homogeneous and nonhomogeneous periodic solutions through all parameters of the system which are spatially homogeneous. To verify our theoretical results, some numerical simulations are also presented. © 2015 Wiley Periodicals, Inc. Complexity 21: 34–43, 2016     

Interactions of Turing and Hopf bifurcations in an additional food provided diffusive predator-prey model

Complex spatiotemporal dynamics of a diffusive predator-prey system involving additional food supply to predator and intra-specific competition among predator, are investigated. We establish critical conditions of the occurrence of Turing instability, which are necessary and sufficient. Furthermore, we also establish conditions of the occurrence of codimension-2 Turing-Hopf bifurcation and Turing-Turing bifurcation, by exploring interactions of Turing bifurcations and Hopf bifurcation. For Turing-Hopf bifurcation, by analyzing normal form truncated to order 3 which are derived by applying normal form method, it is shown that under proper conditions, diffusive predator-prey system generates interesting spatial, temporal and spatiotemporal patterns, including a pair of spatially inhomogeneous steady states, a spatially homogeneous periodic solution and a pair of spatially inhomogeneous periodic solutions. And numerical simulations are also shown to support theory analysis. Moreover, it is found that proper intra-specific competition among predator helps generate complex spatiotemporal dynamics. And, proper additional food supply to predator helps control the population fluctuations of predator and prey, while large quantity and high quality of additional food supply will lead to the extinction of prey and make predator change the source of food, which finally destroy the ecosystem. These investigations might help understand complex spatiotemporal dynamics of predator-prey system involving additional food supply to predator and intra-specific competition among predator, and help conserve species in an ecosystem via supplying suitable additional food.     

Bifurcation analysis on a diffusive Holling–Tanner predator–prey model

This paper is concerned with a diffusive Holling–Tanner predator–prey model subject to homogeneous Neumann boundary condition. By choosing the ratio of intrinsic growth rates of predator to prey λ as the bifurcation parameter, we find that spatially homogeneous and non-homogeneous Hopf bifurcation occur at the positive constant steady state as λ varies. The steady state bifurcation of simple and double eigenvalues are intensively investigated. The techniques of space decomposition and the implicit function theorem are adopted to deal with the case of double eigenvalues. Our results show that this model can exhibit spatially non-homogeneous periodic and stationary patterns induced by the parameter λ. Numerical simulations are presented to illustrate our theoretical results.     

Bifurcation and spatiotemporal patterns in a homogeneous diffusive predator-prey system

A diffusive predator-prey system with Holling type-II predator functional response subject to Neumann boundary conditions is considered. Hopf and steady state bifurcation analysis are carried out in details. In particular we show the existence of multiple spatially non-homogeneous periodic orbits while the system parameters are all spatially homogeneous. Our results and global bifurcation theory also suggest the existence of loops of spatially non-homogeneous periodic orbits and steady state solutions. These results provide theoretical evidences to the complex spatiotemporal dynamics found by numerical simulation.     

Stability and Hopf bifurcation in a diffusive predator–prey system with delay effect

This paper is concerned with a delayed predator–prey system with diffusion effect. First, the stability of the positive equilibrium and the existence of spatially homogeneous and spatially inhomogeneous periodic solutions are investigated by analyzing the distribution of the eigenvalues. Next the direction and the stability of Hopf bifurcation are determined by the normal form theory and the center manifold reduction for partial functional differential equations. Finally, some numerical simulations are carried out for illustrating the theoretical results.     

Bifurcation analysis of a diffusive predator–prey system with a herd behavior and quadratic mortality

In this paper, a diffusive predator–prey system, in which the prey species exhibits herd behavior and the predator species with quadratic mortality, has been studied. The stability of positive constant equilibrium, Hopf bifurcations, and diffusion‐driven Turing instability are investigated under the Neumann boundary condition. The explicit condition for the occurrence of the diffusion‐driven Turing instability is derived, which is determined by the relationship of the diffusion rates of two species. The formulas determining the direction and the stability of Hopf bifurcations depending on the parameters of the system are derived. Finally, numerical simulations are carried out to verify and extend the theoretical results and show the existence of spatially homogeneous periodic solutions and nonconstant steady states. Copyright © 2014 John Wiley & Sons, Ltd.     

Diffusion-driven stability and bifurcation in a predator–prey system with Ivlev-type functional response

A diffusive predator–prey system with Ivlev-type functional response subject to Neumann boundary conditions is considered. Hopf and steady-state bifurcation analysis are carried out in detail. First, the stability of the positive equilibrium and the existence of spatially homogeneous and inhomogeneous periodic solutions are investigated by analysing the distribution of the eigenvalues. The direction and stability of Hopf bifurcation are determined by the normal form theory and the centre manifold reduction for partial functional differential equations and then steady-state bifurcation is studied. Finally, some numerical simulations are carried out for illustrating the theoretical results.     

Bifurcation and Turing pattern formation in a diffusive ratio-dependent predator–prey model with predator harvesting

The ratio-dependent predator–prey model exhibits rich dynamics due to the singularity of the origin. Harvesting in a ratio-dependent predator–prey model is relatively an important research project from both ecological and mathematical points of view. In this paper, we study the temporal, spatial and spatiotemporal dynamics of a ratio-dependent predator–prey diffusive model where the predator population harvest at catch-per-unit-effort hypothesis. For the spatially homogeneous model, we derive conditions for determining the direction of Hopf bifurcation and the stability of the bifurcating periodic solution by the center manifold and the normal form theory. For the reaction–diffusion model, firstly it is shown that Turing (diffusion-driven) instability occurs, which induces spatial inhomogeneous patterns. Then it is demonstrated that the model exhibit Hopf bifurcation which produces temporal inhomogeneous patterns. Finally, the existence and non-existence of positive non-constant steady-state solutions are established. Moreover, numerical simulations are performed to visualize the complex dynamic behavior.     

Multiple bifurcations in a delayed predator–prey diffusion system with a functional response

The present paper is concerned with a delayed predator–prey diffusion system with a Beddington–DeAngelis functional response and homogeneous Neumann boundary conditions. If the positive constant steady state of the corresponding system without delay is stable, by choosing the delay as the bifurcation parameter, we can show that the increase of the delay can not only cause spatially homogeneous Hopf bifurcation at the positive constant steady state but also give rise to spatially heterogeneous ones. In particular, under appropriate conditions, we find that the system has a Bogdanov–Takens singularity at the positive constant steady state, whereas this singularity does not occur for the corresponding system without diffusion. In addition, by applying the normal form theory and center manifold theorem for partial functional differential equations, we give normal forms of Hopf bifurcation and Bogdanov–Takens bifurcation and the explicit formula for determining the properties of spatial Hopf bifurcations.     

Predator invasion in predator–prey model with prey-taxis in spatially heterogeneous environment

We consider a predator–prey model with prey-taxis and Holling-type II functional responses in a spatially heterogeneous environment to analyze the effects of prey-taxis and the heterogeneity of an environment on predator invasion. To achieve our goal, we investigate the stability of semi-trivial solution in which the predator is absent. It is known that both the predator diffusion and the death rate contribute to the predator invasion in a heterogeneous habitat when there is no prey-taxis. In this paper, we show that predator invasion is affected by the prey-taxis and diffusions of the prey-taxis model for a certain range of predator death rates in a heterogeneous environment. Furthermore, in cases where predator invasion by predator diffusion does not occur in a particular death rate range of the predator, predator invasion can occur by prey-taxis in a spatially heterogeneous habitat. In addition, we compare this phenomenon to the corresponding predator–prey model with ratio-dependent functional responses. It is observed that none of the predator’s diffusion and prey-taxis affect the predator’s invasion, and that only the predator’s death rate contributes to predator invasion for the model with ratio-dependent functional responses.     

Stability,Steady‐State Bifurcations,and Turing Patterns in a Predator–Prey Model with Herd Behavior and Prey‐taxis

In this paper, we consider a predator–prey model with herd behavior and prey‐taxis subject to the homogeneous Neumann boundary condition. First, by analyzing the characteristic equation, the local stability of the positive equilibrium is discussed. Then, choosing prey‐tactic sensitivity coefficient as the bifurcation parameter, we obtain a branch of nonconstant solutions bifurcating from the positive equilibrium by an abstract bifurcation theory, and find the stable bifurcating solutions near the bifurcation point under suitable conditions. We have shown that prey‐taxis can destabilize the uniform equilibrium and yields the occurrence of spatial patterns. Furthermore, some numerical simulations to illustrate the theoretical analysis are also carried out, Turing patterns such as spots pattern, spots–strip pattern, strip pattern, stable nonconstant steady‐state solutions, and spatially inhomogeneous periodic solutions are obtained, which also expand our theoretical results.     

A qualitative study on general Gause-type predator–prey models with non-monotonic functional response

In this paper, we study a diffusive predator–prey model with general growth rates and non-monotonic functional response under homogeneous Neumann boundary condition. A local existence of periodic solutions and the asymptotic behavior of spatially inhomogeneous solutions are investigated. Moreover, we show the existence and non-existence of non-constant positive steady-state solutions. Especially, to show the existence of non-constant positive steady-states, the fixed point index theory is used without estimating the lower bounds of positive solutions. More precisely, calculating the indexes at the trivial, semi-trivial and positive constant solutions, some sufficient conditions for the existence of non-constant positive steady-state solutions are studied. This is in contrast to the works in previous papers. Furthermore, on obtaining these results, we can observe that the monotonicity of a prey isocline at the positive constant solution plays an important role.     

Hopf bifurcation analysis in a one-dimensional Schnakenberg reaction-diffusion model

In this paper, we study the Hopf bifurcation phenomenon of a one-dimensional Schnakenberg reaction-diffusion model subject to the Neumann boundary condition. Our results reveal that both spatially homogeneous periodic solutions and spatially heterogeneous periodic solution exist. Moreover, we conclude that the spatially homogeneous periodic solutions are locally asymptotically stable and the spatially heterogeneous periodic solutions are unstable. In addition, we give specific examples to illustrate the phenomenon that coincides with our theoretical results.     

Spatially homogeneous copulas

We consider spatially homogeneous copulas, i.e. copulas whose corresponding measure is invariant under a special transformations of \([0,1]^2\), and we study their main properties with a view to possible use in stochastic models. Specifically, we express any spatially homogeneous copula in terms of a probability measure on [0, 1) via the Markov kernel representation. Moreover, we prove some symmetry properties and demonstrate how spatially homogeneous copulas can be used in order to construct copulas with surprisingly singular properties. Finally, a generalization of spatially homogeneous copulas to the so-called (m, n)-spatially homogeneous copulas is studied and a characterization of this new family of copulas in terms of the Markov \(*\)-product is established.

     

A predator–prey system with stage-structure for predator and nonlocal delay

This paper deals with the behavior of solutions to the reaction–diffusion system under homogeneous Neumann boundary condition, which describes a prey–predator model with nonlocal delay. Sufficient conditions for the global stability of each equilibrium are derived by the Lyapunov functional and the results show that the introduction of stage-structure into predator positively affects the coexistence of prey and predator. Numerical simulations are performed to illustrate the results.     

Positive solutions of a diffusive prey–predator model in a heterogeneous environment

In this paper, we investigate a diffusive prey–predator model in a spatially degenerate heterogeneous environment. We are concerned with the positive solutions of the model, and obtain some results for the existence and non-existence of positive solutions. Moreover, the multiplicity, stability and asymptotical behaviors of positive solutions with respect to the parameters are also studied.     

Hopf bifurcation analysis in the 1-D Lengyel-Epstein reaction-diffusion model

The Lengyel-Epstein model with diffusion and homogeneous Neumann boundary condition is considered in this paper. We give the existence of multiple spatially non-homogeneous periodic solutions though all the parameters of the system are spatially homogeneous.     

Spatiotemporal dynamics analysis and optimal control method for an SI reaction-diffusion propagation model

In the new social media era, it is becoming increasingly important to explore the propagation rules for rumors in social networks. This article is concerned with investigating a diffusive susceptible-infected rumor propagation model with a nonlinear propagation function in a spatially heterogeneous environment. We establish the uniform persistence and analyze the asymptotic behavior of the rumor-spreading steady state for the spatially heterogeneous model when one of the diffusion coefficients tends to zero. Moreover, to better reflect the effect of a time delay on the process of rumor propagation, we establish a spatially homogeneous model with a time delay and prove the existence and local stability of the corresponding equilibrium point. Furthermore, the optimal control in the spatially homogeneous environment case is derived. Finally, several numerical simulations are performed to verify the theoretical results in both spatially heterogeneous and spatially homogeneous systems.