Stability and Hopf bifurcation of a diffusive predator–prey model with predator saturation and competition

This article discusses a predator–prey system with predator saturation and competition functional response. The local stability, existence of a Hopf bifurcation at the coexistence equilibrium and stability of bifurcating periodic solutions are obtained in the absence of diffusion. Further, we discuss the diffusion-driven instability, Hopf bifurcation for corresponding diffusion system with zero flux boundary condition and Turing instability region regarding the parameters are established. Finally, numerical simulations supporting the theoretical analysis are also included.     

Dynamics of a ratio‐dependent stage‐structured predator‐prey model with delay

In this paper, we investigate the dynamics of a time‐delay ratio‐dependent predator‐prey model with stage structure for the predator. This predator‐prey system conforms to the realistically biological environment. The existence and stability of the positive equilibrium are thoroughly analyzed, and the sufficient and necessary conditions for the stability and instability of the positive equilibrium are obtained for the case without delay. Then, the influence of delay on the dynamics of the system is investigated using the geometric criterion developed by Beretta and Kuang. 26 We show that the positive steady state can be destabilized through a Hopf bifurcation and there exist stability switches under some conditions. The formulas determining the direction and the stability of Hopf bifurcations are explicitly derived by using the center manifold reduction and normal form theory. Finally, some numerical simulations are performed to illustrate and expand our theoretical results.     

Stability and cross-diffusion-driven instability in a diffusive predator–prey system with hunting cooperation functional response

This paper presents a qualitative study of a diffusive predator–prey system with the hunting cooperation functional response. For the system without diffusion, the existence, stability and Hopf bifurcation of the positive equilibrium are explicitly determined. It is shown that the hunting cooperation affects not only the existence of the positive equilibrium but also the stability. For the diffusive system, the stability and cross-diffusion driven Turing instability are investigated according to the relationship of the self-diffusion and the cross-diffusion coefficients. Stability and cross-diffusion instability regions are theoretically determined in the plane of the cross-diffusion coefficients. The technique of multiple time scale is employed to deduce the amplitude equation of Turing bifurcation and then pattern dynamics driven by the cross-diffusion is also investigated by the corresponding amplitude equation.     

Turing instability and Hopf bifurcation in a diffusive Leslie–Gower predator–prey model

In this paper, a reaction‐diffusion predator–prey system that incorporates the Holling‐type II and a modified Leslie‐Gower functional responses is considered. For ODE, the local stability of the positive equilibrium is investigated and the specific conditions are obtained. For partial differential equation, we consider the dissipation and persistence of solutions, the Turing instability of the equilibrium solutions, and the Hopf bifurcation. By calculating the normal form, we derive the formulae, which can determine the direction and the stability of Hopf bifurcation according to the original parameters of the system. We also use some numerical simulations to illustrate our theoretical results. Copyright © 2016 John Wiley & Sons, Ltd.     

Effect of herd shape in a diffusive predator-prey model with time delay

In this paper, we deal with the effect of the shape of herd behavior on the interaction between predator and prey. The model analysis was studied in three parts. The first, The analysis of the system in the absence of spatial diffusion and the time delay, where the local stability of the equilibrium states, the existence of Hopf bifurcation have been investigated. For the second part, the spatiotemporal dynamics introduce by self diffusion was determined, where the existence of Hopf bifurcation, Turing driven instability, Turing-Hopf bifurcation point have been proved. Further, the order of Hopf bifurcation points and regions of the stability of the non trivial equilibrium state was given. In the last part of the paper, we studied the delay effect on the stability of the non trivial equilibrium, where we proved that the delay can lead to the instability of interior equilibrium state, and also the existence of Hopf bifurcation. A numerical simulation was carried out to insure the theoretical results.     

Dynamics and pattern formations in diffusive predator‐prey models with two prey‐taxis

A reaction‐diffusion two‐predator‐one‐prey system with prey‐taxis describes the spatial interaction and random movement of predator and prey species, as well as the spatial movement of predators pursuing preys. The global existence and boundedness of solutions of the system in bounded domains of arbitrary spatial dimension and any small prey‐taxis sensitivity coefficient are investigated by the semigroup theory. The spatial pattern formation induced by the prey‐taxis is characterized by the Turing type linear instability of homogeneous state; it is shown that prey‐taxis can both compress and prompt the spatial patterns produced through diffusion‐induced instability in two‐predator‐one‐prey systems.     

The effect of prey refuge and time delay on a diffusive predator‐prey system with hyperbolic mortality

In this article, we investigate the effect of prey refuge and time delay on a diffusive predator‐prey system with Holling II functional response and hyperbolic mortality subject to Neumann boundary condition. More precisely, we study Turing instability of positive equilibrium by using refuge as parameter, instability and Hopf bifurcation induced by time delay. In addition, by the theory of normal form and center manifold, we derive conditions for determining the bifurcation direction and the stability of the bifurcating periodic solution. © 2016 Wiley Periodicals, Inc. Complexity 21: 446–459, 2016     

Consequences of refuge and diffusion in a spatiotemporal predator–prey model

In this investigation, we offer and examine a predator–prey interacting model with prey refuge in proportion to both the species and Beddington–DeAngelis functional response. We first prove the well-posedness of the temporal and spatiotemporal models which are restricted in a positive invariant region. Then for the temporal model, we analyse its temporal dynamics including uniform boundedness, permanence, stability of all feasible non-negative equilibria and show that refugia can induce periodic oscillation via Hopf bifurcation around the unique positive equilibrium; for the spatiotemporal model, we not only investigate its permanence, stability of non-negative constant steady states and Turing instability but also study the existence and non-existence of non-constant positive steady states by Leray–Schauder degree theory. The key observation is that the coefficient of refuge cooperates a significant part in modifying the dynamics of the current system and mediates the population permanence, stability of coexisting equilibrium and even the Turing instability parameter space. Finally, general numerical simulation consequences are given to illustrate the validity of the theoretical results. Through numerical simulations, one observes that the model dynamics shows prey refugia and self-diffusion control spatiotemporal pattern growth to spots, stripe–spot mixtures and stripes reproduction. The outcomes assign that the dynamics of the model with prey refuge is not simple, but rich and complex. Additionally, numerical simulations show that the other model parameters have an important effect on species’ spatially inhomogeneous distribution, which results in the formation of spots pattern, mixture of spots and stripes pattern, mixture of spots, stripes and rings pattern and anti-spot pattern. This may improve the model dynamics of the prey refuge on the reaction–diffusion predator–prey system.     

Stability and instability analysis for a ratio-dependent predator–prey system with diffusion effect

In this paper, we consider a ratio-dependent predator–prey system with diffusion. And we mainly discuss the following problems: (1) stability and Hopf bifurcation analysis of the positive equilibrium for the reduced ODE system; (2) Diffusion-driven instability of the equilibrium solution; (3) Hopf bifurcations for the corresponding diffusion system with homogeneous Neumann boundary conditions. In order to verify our theoretical results, some numerical simulations are also included, respectively.     

Dynamics in a diffusive predator–prey system with a constant prey refuge and delay

In this paper, a diffusive predator–prey system with a constant prey refuge and time delay subject to Neumann boundary condition is considered. Local stability and Turing instability of the positive equilibrium are studied. The effect of time delay on the model is also obtained, including locally asymptotical stability and existence of Hopf bifurcation at the positive equilibrium. And the properties of Hopf bifurcation are determined by center manifold theorem and normal form theorem of partial functional differential equations. Some numerical simulations are carried out.     

Interactions of Turing and Hopf bifurcations in an additional food provided diffusive predator-prey model

Complex spatiotemporal dynamics of a diffusive predator-prey system involving additional food supply to predator and intra-specific competition among predator, are investigated. We establish critical conditions of the occurrence of Turing instability, which are necessary and sufficient. Furthermore, we also establish conditions of the occurrence of codimension-2 Turing-Hopf bifurcation and Turing-Turing bifurcation, by exploring interactions of Turing bifurcations and Hopf bifurcation. For Turing-Hopf bifurcation, by analyzing normal form truncated to order 3 which are derived by applying normal form method, it is shown that under proper conditions, diffusive predator-prey system generates interesting spatial, temporal and spatiotemporal patterns, including a pair of spatially inhomogeneous steady states, a spatially homogeneous periodic solution and a pair of spatially inhomogeneous periodic solutions. And numerical simulations are also shown to support theory analysis. Moreover, it is found that proper intra-specific competition among predator helps generate complex spatiotemporal dynamics. And, proper additional food supply to predator helps control the population fluctuations of predator and prey, while large quantity and high quality of additional food supply will lead to the extinction of prey and make predator change the source of food, which finally destroy the ecosystem. These investigations might help understand complex spatiotemporal dynamics of predator-prey system involving additional food supply to predator and intra-specific competition among predator, and help conserve species in an ecosystem via supplying suitable additional food.     

Investigating the Turing Conditions for Diffusion-driven Instability in Predator-prey System with Hunting Cooperation Functional Response

In this paper, we focus on stability analysis of steady-state solutions of a predator-prey system with hunting cooperation functional response. The results show that the Turing instability can be affected not only the existence of hunting cooperation, but also the diffusion coefficients: (1) in the absence of predator diffusion, diffusion-driven instability can be induced by hunting cooperation, but no stable patterns appear; (2) the system can occur diffusion-driven instability and Turing patterns, when both predator and prey have diffusion, and the diffusion coefficient of prey is greater than that of the predator. The numerical simulations of two cases are presented to verify the validity of our theoretical results.     

Bifurcation and Turing pattern formation in a diffusive ratio-dependent predator–prey model with predator harvesting

The ratio-dependent predator–prey model exhibits rich dynamics due to the singularity of the origin. Harvesting in a ratio-dependent predator–prey model is relatively an important research project from both ecological and mathematical points of view. In this paper, we study the temporal, spatial and spatiotemporal dynamics of a ratio-dependent predator–prey diffusive model where the predator population harvest at catch-per-unit-effort hypothesis. For the spatially homogeneous model, we derive conditions for determining the direction of Hopf bifurcation and the stability of the bifurcating periodic solution by the center manifold and the normal form theory. For the reaction–diffusion model, firstly it is shown that Turing (diffusion-driven) instability occurs, which induces spatial inhomogeneous patterns. Then it is demonstrated that the model exhibit Hopf bifurcation which produces temporal inhomogeneous patterns. Finally, the existence and non-existence of positive non-constant steady-state solutions are established. Moreover, numerical simulations are performed to visualize the complex dynamic behavior.     

Bifurcation analysis of a diffusive predator–prey system with nonconstant death rate and Holling III functional response

In this paper, a diffusive predator–prey system with Holling III functional response and nonconstant death rate subject to Neumann boundary condition is considered. We study the stability of equilibria, and Turing instability of the positive equilibrium. We also perform a detailed Hopf bifurcation analysis to PDE system, and derive conditions for determining the bifurcation direction and the stability of the bifurcating periodic solution. In addition, some numerical simulations are carried out.     

Herd behavior in a predator–prey model with spatial diffusion: bifurcation analysis and Turing instability

We consider in this paper an ecological model, in a predator–prey interaction with the presence of a herd behavior. For the analysis of the model, the existence of positive solution and also the existence Hopf bifurcation, Turing driven instability, and Turing–Hopf bifurcation point have bee proved. Then by calculating the normal form, on the center of the manifold associated to the Hopf bifurcation points, the stability of the periodic solution has been proved. In the last part of the paper, numerical simulations has been given to illustrate our theoretical analysis.     

Spatiotemporal dynamics in a predator-prey model with a functional response increasing in both predator and prey densities

In this paper, we studied a diffusive predator-prey model with a functional response increasing in both predator and prey densities. The Turing instability and local stability are studied by analyzing the eigenvalue spectrum. Delay induced Hopf bifurcation is investigated by using time delay as bifurcation parameter. Some conditions for determining the property of Hopf bifurcation are obtained by utilizing the normal form method and center manifold reduction for partial functional differential equation.     

Turing instability for a ratio-dependent predator-prey model with diffusion

Ratio-dependent predator-prey models have been increasingly favored by field ecologists where predator-prey interactions have to be taken into account the process of predation search. In this paper we study the conditions of the existence and stability properties of the equilibrium solutions in a reaction-diffusion model in which predator mortality is neither a constant nor an unbounded function, but it is increasing with the predator abundance. We show that analytically at a certain critical value a diffusion driven (Turing type) instability occurs, i.e. the stationary solution stays stable with respect to the kinetic system (the system without diffusion). We also show that the stationary solution becomes unstable with respect to the system with diffusion and that Turing bifurcation takes place: a spatially non-homogenous (non-constant) solution (structure or pattern) arises. A numerical scheme that preserve the positivity of the numerical solutions and the boundedness of prey solution will be presented. Numerical examples are also included.     

Spatiotemporal dynamics of reaction–diffusion models of interacting populations

The present investigation deals with the necessary conditions for Turing instability with zero-flux boundary conditions that arise in a ratio-dependent predator–prey model involving the influence of logistic population growth in prey and intra-specific competition among predators described by a system of non-linear partial differential equations. The prime objective is to investigate the parametric space for which Turing spatial structure takes place and to perform extensive numerical simulation from both the mathematical and the biological points of view in order to examine the role of diffusion coefficients in Turing instability. Various spatiotemporal distributions of interacting species through Turing instability in two dimensional spatial domain are portrayed and analyzed at length in order to substantiate the applicability of the present model.     

Turing patterns induced by cross-diffusion in a predator-prey system in presence of habitat complexity

In this paper, we have investigated the phenomena of Turing pattern formation in a predator-prey model with habitat complexity in presence of cross diffusion. Using the linear stability analysis, the conditions for the existence of stationary pattern and the existence of Hopf bifurcation are obtained. It is shown analytically that the presence of cross diffusion in the system supports the formation of Turing pattern. Two parameter bifurcation analysis are done analytically and corresponding bifurcation diagrams are presented numerically. A series of simulation results are plotted for different biologically meaningful parameter values. Effects of variation of habitat complexity and the predator mortality rate and birth rate of prey on pattern formation are also reported. It is shown that cross-diffusion can lead to a wide variety of spatial and spatiotemporal pattern formation. It is found that the model exhibits spot and stripe pattern, and coexistence of both spot and strip patterns under the zero flux boundary condition. It is observed that cross-diffusion, habitat complexity, birth rate of prey and predator’s mortality rate play a significant role in the pattern formation of a distributed population system of predator-prey type.     

Spatial Dynamics of a Diffusive Predator-prey Model with Leslie-Gower Functional Response and Strong Allee Effect

In this paper, spatial dynamics of a diffusive predator-prey model with Leslie-Gower functional response and strong Allee effect is studied. Firstly, we obtain the critical condition of Hopf bifurcation and Turing bifurcation of the PDE model. Secondly, taking self-diffusion coefficient of the prey as bi- furcation parameter, the amplitude equations are derived by using multi-scale analysis methods. Finally, numerical simulations are carried out to verify our theoretical results. The simulations show that with the decrease of self- diffusion coefficient of the prey, the preys present three pattern structures: spot pattern, mixed pattern, and stripe pattern. We also observe the transi- tion from spot patterns to stripe patterns of the prey by changing the intrinsic growth rate of the predator. Our results reveal that both diffusion and the intrinsic growth rate play important roles in the spatial distribution of species.