Dynamical analysis of a delayed singular prey–predator economic model with stochastic fluctuations

This article studies a delayed singular prey–predator economic model with stochastic fluctuations, which is described by differential‐algebraic equations due to a economic theory. Local stability and Hopf bifurcation condition are described on the delayed singular prey–predator economic model within deterministic environment. It reveals the sensitivity of the model dynamics on gestation time delay. A phenomenon of Hopf bifurcation occurs as the gestation time delay increases through a certain threshold. Subsequently, a singular stochastic prey–predator economic model with time delay is obtained by introducing Gaussian white noise terms to the above deterministic model system. The fluctuation intensity of population and harvest effort are calculated by Fourier transforms method. Numerical simulations are carried out to substantiate these theory analysis. © 2013 Wiley Periodicals, Inc. Complexity 19: 23–29, 2014     

Novel dynamics of a predator–prey system with harvesting of the predator guided by its population

A predator–prey model was extended to include nonlinear harvesting of the predator guided by its population, such that harvesting is only implemented if the predator population exceeds an economic threshold. The proposed model is a nonsmooth dynamic system with switches between the original predator-prey model (free subsystem) and a model with nonlinear harvesting (harvesting subsystem). We initially examine the dynamics of both the free and the harvesting subsystems, and then we investigate the dynamics of the switching system using theories of nonsmooth systems. Theoretical results showed that the harvesting subsystem undergoes multiple bifurcations, including saddle-node, supercritical Hopf, Bogdanov–Takens and homoclinic bifurcations. The switching system not only retains all of the complex dynamics of the harvesting system but also exhibits much richer dynamics such as a sliding equilibrium, sliding cycle, boundary node (saddle point) bifurcation, boundary saddle-node bifurcation and buckling bifurcation. Both theoretical and numerical results showed that, by implementing predator population guided harvesting, the predator and prey population could coexist in more scenarios than those in which the predator may go extinct for the continuous harvesting regime. They could either stabilize at an equilibrium or oscillate periodically depending on the value of the economic threshold and the initial value of the system.     

A bioeconomic differential algebraic predator–prey model with nonlinear prey harvesting

In this paper, we propose a bioeconomic differential algebraic predator–prey model with Holling type II functional response and nonlinear prey harvesting. As the nonlinear prey harvesting is introduced, the proposed model displays a complex dynamics in the predator–prey plane. Taking into account of the economic factor, our predator–prey system is established by bioeconomic differential algebraic equations. The effect of economic profit on the proposed model is analyzed by viewing it as a bifurcation parameter. By jointly using the normal form of differential algebraic models and the bifurcation theory, the stability and bifurcations (singularity induced bifurcation, Hopf bifurcation) are discussed. These results obtained here reveal richer dynamics of the bioeconomic differential algebraic predator–prey model with nonlinear prey harvesting, and suggest a guidance for harvesting in the practical word. Finally, numerical simulations are given to demonstrate the results.     

Dynamics analysis and numerical simulations of a stochastic non-autonomous predator–prey system with impulsive effects

In this paper, we propose a stochastic non-autonomous Lotka–Volterra predator–prey model with impulsive effects and investigate its stochastic dynamics. We first prove that the subsystem of the system has a unique periodic solution which is globally attractive. Furthermore, we obtain the threshold value in the mean which governs the stochastic persistence and the extinction of the prey–predator system. Our results show that the stochastic noises and impulsive perturbations have crucial effects on the persistence and extinction of each species. Finally, we use the different stochastic noises and impulsive effects parameters to provide a series of numerical simulations to illustrate the analytical results.     

Bifurcation and Turing pattern formation in a diffusive ratio-dependent predator–prey model with predator harvesting

The ratio-dependent predator–prey model exhibits rich dynamics due to the singularity of the origin. Harvesting in a ratio-dependent predator–prey model is relatively an important research project from both ecological and mathematical points of view. In this paper, we study the temporal, spatial and spatiotemporal dynamics of a ratio-dependent predator–prey diffusive model where the predator population harvest at catch-per-unit-effort hypothesis. For the spatially homogeneous model, we derive conditions for determining the direction of Hopf bifurcation and the stability of the bifurcating periodic solution by the center manifold and the normal form theory. For the reaction–diffusion model, firstly it is shown that Turing (diffusion-driven) instability occurs, which induces spatial inhomogeneous patterns. Then it is demonstrated that the model exhibit Hopf bifurcation which produces temporal inhomogeneous patterns. Finally, the existence and non-existence of positive non-constant steady-state solutions are established. Moreover, numerical simulations are performed to visualize the complex dynamic behavior.     

Global bifurcation of solutions for a predator–prey model with prey‐taxis

We study pattern formations in a predator–prey model with prey‐taxis. It is proved that a branch of nonconstant solutions can bifurcate from the positive equilibrium only when the chemotactic is repulsive. Furthermore, we find the stable bifurcating solutions near the bifurcation point under suitable conditions. Copyright © 2014 John Wiley & Sons, Ltd.     

A delayed prey–predator model with Crowley–Martin‐type functional response including prey refuge

In this paper, we have studied a prey–predator model living in a habitat that divided into two regions: an unreserved region and a reserved (refuge) region. The migration between these two regions is allowed. The interaction between unreserved prey and predator is Crowley–Martin‐type functional response. The local and global stability of the system is discussed. Further, the system is extended to incorporate the effect of time delay. Then the dynamical behavior of the system is analyzed, taking delay as a bifurcation parameter. The direction of Hopf bifurcation and the stability of the bifurcated periodic solution are determined with the help of normal form theory and centre manifold theorem. We have also discussed the influence of prey refuge on densities of prey and predator species. The analytical results are supplemented with numerical simulations. Copyright © 2017 John Wiley & Sons, Ltd.     

A discrete-time model with non-monotonic functional response and strong Allee effect in prey

In this paper, we investigate the impact of strong Allee effect on the stability of a discrete-time predator–prey model with a non-monotonic functional response. The dynamics of discrete-time predator–prey models with strong Allee effect is studied earlier. But, the mathematical investigations of predator–prey dynamics in discrete-time set up with Holling type-IV functional response and strong Allee effect in prey are lacking. The proposed model supports the coexistence of two steady states, and the mathematical features of the model are analyzed based on local stability and bifurcation theory. By considering the Allee parameter as the bifurcation parameter, we provide sufficient conditions for the flip and the Neimark–Sacker bifurcations. We observe that Allee parameter plays a significant role in the dynamics of the system.     

A stage-structured food chain model with stage dependent predation: Existence of codimension one and codimension two bifurcations

Stage-structured predator–prey models exhibit rich and interesting dynamics compared to homogeneous population models. The objective of this paper is to study the bifurcation behavior of stage-structured prey–predator models that admit stage-restricted predation. It is shown that the model with juvenile-only predation exhibits Hopf bifurcation with the growth rate of the adult prey as the bifurcation parameter; also, depending on parameter values, a stable limit cycle will emerge, that is, the bifurcation will be of supercritical nature. On the other hand, the analysis of the model with adult-stage predation shows that the system admits a fold-Hopf bifurcation with the adult growth rate and the predator mortality rate as the two bifurcation parameters. We also demonstrate the existence of a unique limit cycle arising from this codimension-2 bifurcation. These results reveal far richer dynamics compared to models without stage-structure. Numerical simulations are done to support analytical results.     

A Leslie–Gower predator–prey model with disease in prey incorporating a prey refuge

In this paper, a predator–prey Leslie–Gower model with disease in prey has been developed. The total population has been divided into three classes, namely susceptible prey, infected prey and predator population. We have also incorporated an infected prey refuge in the model. We have studied the positivity and boundedness of the solutions of the system and analyzed the existence of various equilibrium points and stability of the system at those equilibrium points. We have also discussed the influence of the infected prey refuge on each population density. It is observed that a Hopf bifurcation may occur about the interior equilibrium taking refuge parameter as bifurcation parameter. Our analytical findings are illustrated through computer simulation using MATLAB, which show the reliability of our model from the eco-epidemiological point of view.     

Stability of a predator–prey model with refuge effect

We consider a predator–prey model, where some prey are completely free from predation within a temporal or spacial refuge. The most common type of spacial refuge, that we investigate here, takes the form where a constant proportion of the prey population is protected. The model is a modification of the classical Nicholson–Bailey host-parasitoid model. In this paper, we study the effect of the presence of refuge on the stability and bifurcation of the system. Moreover, we provide a detailed analysis of the Neimark–Sacker bifurcation of the model.     

Stationary distribution and extinction of a stochastic predator–prey model with distributed delay

In this paper, we focus on a stochastic predator–prey model with distributed delay. We first obtain the existence of a stationary distribution to the positive solutions by stochastic Lyapunov function method. Then we establish sufficient conditions for extinction of the predator population, that is, the prey population is survival and the predator population is extinct.     

Bifurcation analysis of a predator–prey model with memory-based diffusion

In this paper, we investigate the predator–prey model equipped with Fickian diffusion and memory-based diffusion of predators. The stability and bifurcation analysis explores the impacts of the memory-based diffusion and the averaged memory period on the dynamics near the positive steady state. Specifically, when the memory-based diffusion coefficient is less than a critical value, we show that the stability of the positive steady state can be destabilized as the average memory period increases, which leads to the occurrence of Hopf bifurcations. Moreover, we also analyze the bifurcation properties using the central manifold theorem and normal form theory. This allows us to prove the existence of stable spatially inhomogeneous periodic solutions arising from Hopf bifurcation. In addition, the sufficient and necessary conditions for the occurrence of stability switches are also provided.     

Delay-induced Bogdanov–Takens bifurcation in a Leslie–Gower predator–prey model with nonmonotonic functional response

This work is concerned with the dynamics of a Leslie–Gower predator–prey model with nonmonotonic functional response near the Bogdanov–Takens bifurcation point. By analyzing the characteristic equation associated with the nonhyperbolic equilibrium, the critical value of the delay inducing the Bogdanov–Takens bifurcation is obtained. In this case, the dynamics near this nonhyperbolic equilibrium can be reduced to the study of the dynamics of the corresponding normal form restricted to the associated two-dimensional center manifold. The bifurcation diagram near the Bogdanov–Takens bifurcation point is drawn according to the obtained normal form. We show that the change of delay can result in heteroclinic orbit, homoclinic orbit and unstable limit cycle.     

Stability,Steady‐State Bifurcations,and Turing Patterns in a Predator–Prey Model with Herd Behavior and Prey‐taxis

In this paper, we consider a predator–prey model with herd behavior and prey‐taxis subject to the homogeneous Neumann boundary condition. First, by analyzing the characteristic equation, the local stability of the positive equilibrium is discussed. Then, choosing prey‐tactic sensitivity coefficient as the bifurcation parameter, we obtain a branch of nonconstant solutions bifurcating from the positive equilibrium by an abstract bifurcation theory, and find the stable bifurcating solutions near the bifurcation point under suitable conditions. We have shown that prey‐taxis can destabilize the uniform equilibrium and yields the occurrence of spatial patterns. Furthermore, some numerical simulations to illustrate the theoretical analysis are also carried out, Turing patterns such as spots pattern, spots–strip pattern, strip pattern, stable nonconstant steady‐state solutions, and spatially inhomogeneous periodic solutions are obtained, which also expand our theoretical results.     

Bifurcation analysis in a discrete differential-algebraic predator–prey system

The discrete-time predator–prey biological economic system obtained by Euler method is investigated. Some conditions for the system to undergo flip bifurcation and Neimark–Sacker bifurcation are derived by using new normal form of differential-algebraic system, center mainfold theorem and bifurcation theory. Numerical simulations are given to show the effectiveness of our results and also to exhibit period-doubling bifurcation in orbits of period 2, 4, 8 and chaotic sets. The results obtained here reveal far richer dynamics in discrete differential-algebraic biological economic system. The contents are interesting in mathematics and biology.     

Bifurcations in a Diffusive Predator–Prey Model with Beddington–DeAngelis Functional Response and Nonselective Harvesting

In this paper, we discuss the dynamics of a predator–prey model with Beddington–DeAngelis functional response and nonselective harvesting. By using the Lyapunov–Schmidt reduction, we obtain the existence of spatially nonhomogeneous steady-state solution. The stability and existence of Hopf bifurcation at the spatially nonhomogeneous steady-state solution with the change of a specific parameter are investigated by analyzing the distribution of the eigenvalues. We also get an algorithm for determining the bifurcation direction of the Hopf bifurcating periodic solutions near the nonhomogeneous steady-state solution. Finally, we show some numerical simulations to verify our analytical results.

     

Dynamics of a Stochastic Predator–Prey Model with Stage Structure for Predator and Holling Type II Functional Response

In this paper, we develop and study a stochastic predator–prey model with stage structure for predator and Holling type II functional response. First of all, by constructing a suitable stochastic Lyapunov function, we establish sufficient conditions for the existence and uniqueness of an ergodic stationary distribution of the positive solutions to the model. Then, we obtain sufficient conditions for extinction of the predator populations in two cases, that is, the first case is that the prey population survival and the predator populations extinction; the second case is that all the prey and predator populations extinction. The existence of a stationary distribution implies stochastic weak stability. Numerical simulations are carried out to demonstrate the analytical results.     

Bogdanov–Takens bifurcation in a predator–prey model

In this paper, we investigate a class of predator–prey model with age structure and discuss whether the model can undergo Bogdanov–Takens bifurcation. The analysis is based on the normal form theory and the center manifold theory for semilinear equations with non-dense domain combined with integrated semigroup theory. Qualitative analysis indicates that there exist some parameter values such that this predator–prey model has an unique positive equilibrium which is Bogdanov–Takens singularity. Moreover, it is shown that under suitable small perturbation, the system undergoes the Bogdanov–Takens bifurcation in a small neighborhood of this positive equilibrium.     

Bifurcation and complexity of Monod type predator–prey system in a pulsed chemostat

In this paper, we introduce and study a model of a predator–prey system with Monod type functional response under periodic pulsed chemostat conditions, which contains with predator, prey, and periodically pulsed substrate. We investigate the subsystem with substrate and prey and study the stability of the periodic solutions, which are the boundary periodic solutions of the system. The stability analysis of the boundary periodic solution yields an invasion threshold. By use of standard techniques of bifurcation theory, we prove that above this threshold there are periodic oscillations in substrate, prey and predator. Simple cycles may give way to chaos in a cascade of period-doubling bifurcations. Furthermore, by comparing bifurcation diagrams with different bifurcation parameters, we can see that the impulsive system shows two kinds of bifurcations, whose are period-doubling and period-halfing.