Chaos in periodically forced Holling type IV predator–prey system with impulsive perturbations

The effect of periodic forcing and impulsive perturbations on predator–prey model with Holling type IV functional response is investigated. The periodic forcing is affected by assuming a periodic variation in the intrinsic growth rate of the prey. The impulsive perturbations are affected by introducing periodic constant impulsive immigration of predator. The dynamical behavior of the system is simulated and bifurcation diagrams are obtained for different parameters. The results show that periodic forcing and impulsive perturbation can easily give rise to complex dynamics, including (1) quasi-periodic oscillating, (2) period doubling cascade, (3) chaos, (4) period halfing cascade.     

The dynamical behaviors of a food chain model with impulsive effect and Ivlev functional response

In this paper, a food chain model with Ivlev functional response and impulsive effect of top predator is investigated. Conditions for extinction of mid-level predator are given. By using the Floquet theory of linear τ-period impulsive differential equation and small amplitude perturbation skills, we show that the lowest-level prey and the mid-level predator extinction periodic solution is unstable, while the mid-level predator eradication periodic solution is stable, and meanwhile, we prove that the system is permanent if the impulsive period is larger than some critical value. Furthermore, influences of the impulsive perturbation on the inherent oscillation are studied numerically, which displays complicated behavior including a sequence of direct and inverse cascade of period doubling, period halfing as well as chaos.     

The dynamic complexity of a three-species Beddington-type food chain with impulsive control strategy

In this paper, by using theories and methods of ecology and ordinary differential equation, the dynamics complexity of a prey–predator system with Beddington-type functional response and impulsive control strategy is established. Conditions for the system to be extinct are given by using the Floquet theory of impulsive equation and small amplitude perturbation skills. Furthermore, by using the method of numerical simulation with the international software Maple, the influence of the impulsive perturbations on the inherent oscillation is investigated, which shows rich dynamics, such as quasi-periodic oscillation, narrow periodic window, wide periodic window, chaotic bands, period doubling bifurcation, symmetry-breaking pitchfork bifurcation, period-halving bifurcation and crises, etc. The numerical results indicate that computer simulation is a useful method for studying the complex dynamic systems.     

The study of predator–prey system with defensive ability of prey and impulsive perturbations on the predator

Predator–prey system with non-monotonic functional response and impulsive perturbations on the predator is established. By using Floquet theorem and small amplitude perturbation skills, a locally asymptotically stable prey-eradication periodic solution is obtained when the impulsive period is less than the critical value. Otherwise, if the impulsive period is larger than the critical value, the system is permanent. Further, using numerical simulation method the influences of the impulsive perturbations on the inherent oscillation are investigated. With the increasing of the impulsive value, the system displays a series of complex phenomena, which include (1) quasi-periodic oscillating, (2) period-doubling, (3) period-halfing, (4) non-unique dynamics (meaning that several attractors coexist), (5) attractor crisis and (6) chaotic bands with periodic windows.     

Dynamic complexities of a food chain model with impulsive perturbations and Beddington–DeAngelis functional response

In this paper, the dynamic complexities of a three trophic level food chain system with impulsive perturbations and Beddington–DeAngelis functional responses is studied. Conditions for extinction of predator are given. By using the Floquet theory of impulsive equation and small amplitude perturbation skills, we consider the local stability of predator eradication periodic solution. Further, influences of the impulsive perturbation on the inherent oscillation are studied numerically, which shows the rich dynamics in the positive octant.     

The dynamics of a Beddington-type system with impulsive control strategy

In this paper, by using the theories and methods of ecology and ordinary differential equation, a prey–predator system with Beddington-type functional response and impulsive control strategy is established. Conditions for the system to be extinct are given by using the theories of impulsive equation and small amplitude perturbation skills. It is proved that the system is permanent via the method of comparison involving multiple Liapunov functions. Furthermore, by using the method of numerical simulation, the influence of the impulsive control strategy on the inherent oscillation are investigated, which shows rich dynamics, such as period doubling bifurcation, crises, symmetry-breaking pitchfork bifurcations, chaotic bands, quasi-periodic oscillation, narrow periodic window, wide periodic window, period-halving bifurcation, etc. That will be useful for study of the dynamic complexity of ecosystems.     

Permanence and stability of an Ivlev-type predator–prey system with impulsive control strategies

In this paper, we study a predator–prey system with an Ivlev-type functional response and impulsive control strategies containing a biological control (periodic impulsive immigration of the predator) and a chemical control (periodic pesticide spraying) with the same period, but not simultaneously. We find conditions for the local stability of the prey-free periodic solution by applying the Floquet theory of an impulsive differential equation and small amplitude perturbation techniques to the system. In addition, it is shown that the system is permanent under some conditions by using comparison results of impulsive differential inequalities. Moreover, we add a forcing term into the prey population’s intrinsic growth rate and find the conditions for the stability and for the permanence of this system.     

具有Holling Ⅲ功能反应和阶段结构的Gompertz生态系统的持久性

以生态学与微分方程的理论和方法为基础,建立了一类具有HollingⅢ功能反应和阶段结构的生态Gompertz模型.利用频闪映射,获得了捕食者灭绝周期解,分析了此周期解的全局吸引性.在对食饵进行脉冲收获和捕食者具有成长期时滞条件下,运用脉冲微分方程比较定理和小振幅扰动技巧,获得了系统一致持续生存的条件.     

Mathematics analysis and chaos in an ecological model with an impulsive control strategy

In this paper, on the basis of the theories and methods of ecology and ordinary differential equation, an ecological model with an impulsive control strategy is established. By using the theories of impulsive equation, small amplitude perturbation skills and comparison technique, we get the condition which guarantees the global asymptotical stability of the lowest-level prey and mid-level predator eradication periodic solution. It is proved that the system is permanent. Further, influences of the impulsive perturbation on the inherent oscillation are studied numerically, which shows rich dynamics, such as period-doubling bifurcation, period-halving bifurcation, chaotic band, narrow or wide periodic window, chaotic crises,etc. Moreover, the computation of the largest Lyapunov exponent demonstrates the chaotic dynamic behavior of the model. At the same time, we investigate the qualitative nature of strange attractor by using Fourier spectra. All these results may be useful for study of the dynamic complexity of ecosystems.     

Dynamics of an impulsive control system which prey species share a common predator

In an ecosystem multiple prey species often share a common predator and the interactions between the preys are neutral. In view of these facts and based on a multiple species prey–predator system with Holling IV and II functional responses, an impulsive differential equation to model the process of periodically releasing natural enemies and spraying pesticides at different fixed times for pest control is proposed and investigated. It is proved that there exists a locally asymptotically stable pest-eradication periodic solution under the assumption that the impulsive period is less than some critical value (or the release amount of the predator is greater than another critical value). Permanence conditions are established when the impulsive period is greater than another critical value (or the release amount of the predator is less than some critical value). Numerical results show that the system we consider has more complex dynamics including period solution, quasi-periodic oscillation, chaos, intermittency and crises.     

Analysis of a predator–prey model with Holling II functional response concerning impulsive control strategy

According to biological and chemical control strategy for pest control, we investigate the dynamic behavior of a Holling II functional response predator–prey system concerning impulsive control strategy-periodic releasing natural enemies and spraying pesticide at different fixed times. By using Floquet theorem and small amplitude perturbation method, we prove that there exists a stable pest-eradication periodic solution when the impulsive period is less than some critical value. Further, the condition for the permanence of the system is also given. Numerical results show that the system we consider can take on various kinds of periodic fluctuations and several types of attractor coexistence and is dominated by periodic, quasiperiodic and chaotic solutions, which implies that the presence of pulses makes the dynamic behavior more complex. Finally, we conclude that our impulsive control strategy is more effective than the classical one if we take chemical control efficiently.     

An impulsive controlled eco-epidemic model with disease in the prey

In this paper, we investigate the dynamic behavior of an eco-epidemic model with impulsive control strategy. By using Floquet theorem of impulsive differential equation, we show there is a globally stable prey eradication periodic solution when the impulsive period is less than some critical value. We study the permanence of the system. Numerical simulations show that the complex dynamics of the system depends on the values of impulsive period and impulsive perturbation, for example double period, triple period solutions.     

具有非单调功能反应和脉冲扰动的捕食系统的研究

研究捕食者具有非单调功能反应和周期脉冲扰动的捕食者一食饵系统，利用脉冲微分方程的Floquet理论和比较定理，得到了系统灭绝和持续生存的充分条件．最后，通过数值模拟阐明系统在周期脉冲扰动下的复杂性．     

Analysis of mathematics and dynamics in a food web system with impulsive perturbations and distributed time delay

In this paper, on the basis of the theories and methods of ecology and ordinary differential equation, a food web system with impulsive perturbations and distributed time delay is established. By using the theories of impulsive equation, small amplitude perturbation skills and comparison technique, we get the condition which guarantees the global asymptotical stability of the prey and intermediate predator eradication periodic solution. On this basis, we get that the food web system is permanent if some parameters are satisfied with certain conditions. In order to show that these conditions are effective, the influences of impulsive perturbations on the inherent oscillation and distributed time delay are studied numerically; these show rich dynamics, such as period-halving bifurcation, chaotic band, narrow or wide periodic window, chaotic crises. Moreover, the computation of the largest Lyapunov exponent shows the chaotic dynamic behavior of the model. Meanwhile, we investigate the qualitative nature of strange attractor by using Fourier spectra. All of these results may be useful in the study of the dynamic complexity of ecosystems.     

A study of predator–prey models with the Beddington–DeAnglis functional response and impulsive effect

In this paper, the predator–prey system with the Beddington–DeAngelis functional response is developed, by introducing a proportional periodic impulsive catching or poisoning for the prey populations and a constant periodic releasing for the predator. The Beddington–DeAngelis functional response is similar to the Holling type II functional response but contains an extra term describing mutual interference by predators. This model has the potential to protect predator from extinction, but under some conditions may also lead to extinction of the prey. That is, the system exists a locally stable prey-eradication periodic solution when the impulsive period satisfies an inequality. The condition for permanence is established via the method of comparison involving multiple Liapunov̀ functions. Further, by numerical simulation method the influences of the impulsive perturbations and mutual interference by predators on the inherent oscillation are investigated. With the increasing of releasing for the predator, the system appears a series of complex phenomenon, which include (1) period-doubling, (2) chaos attractor, (3) period-halfing. (4) non-unique dynamics (meaning that several attractors coexist).     

Extinction and permanence of a two-prey two-predator system with impulsive on the predator

In this paper, the dynamic behaviors of a two-prey two-predator system with impulsive effect on the predator of fixed moment are investigated. By applying the Floquet theory of liner periodic impulsive equation, we show that there exists a globally asymptotically stable two-prey eradication periodic solution when the impulsive period is less than some critical value. Further, we prove that the system is permanent if the impulsive period is large than some critical value, and meanwhile the conditions for the extinction of one of the two prey and permanence of the remaining three species are given. Finally, numerical simulation shows that there exists a stable positive periodic solution with a maximum value no larger than a given level. Thus, we can use the stability of the positive periodic solution and its period to control insect pests at acceptably low levels.     

Complex dynamics of a delayed stage-structured predator-prey model with impulsive effect

In this paper, we investigate a stage-structured predator-prey model with periodic harvesting (catching or poisoning) for the prey and stage structure for the predator with constant maturation time delay. Sufficient conditions which guaranteed the global attractivity of predator-extinction periodic solution and permanence of the system are obtained. Furthermore, influences of the impulsive perturbation on the inherent oscillation are studied, which exhibits a wide variety of dynamic behaviors by numerical simulations.     

The study of a predator–prey system with group defense and impulsive control strategy

A predator–prey system with group defense and impulsive control strategy is established. By using Floquet theorem and small amplitude perturbation skills, a locally asymptotically stable prey-eradication periodic solution is obtained when the impulsive period is less than some critical value. Otherwise, if the impulsive period is larger than the critical value, the system is permanent. By using bifurcation theory, we show the existence and stability of positive periodic solution when the pest-eradication lost its stability. Further, numerical examples show that the system considered has more complicated dynamics, such as: (1) quasi-periodic oscillating, (2) period-doubling bifurcation, (3) period-halving bifurcation, (4) non-unique dynamics (meaning that several attractors coexist), (5) attractor crisis, etc. Finally, the biological implications of the results and the impulsive control strategy are discussed.     

Impulsive perturbations in a predator–prey model with dormancy of predators

In this paper, a predator–prey model consisting of active and dormant states of predators with impulsive control strategy is established. Using Floquet theories, the small amplitude perturbation technique and the piecewise Lyapunov function method, the conditions of local and global asymptotical orbital stability of the prey-eradication periodic solution are obtained. The boundness and permanence of the impulsive system are proved by the comparison principle. Through numerical simulations, the effects of the impulsive perturbation on the inherent oscillation are investigated, which implies that the impulsive perturbation can lead to period-doubling bifurcation, chaos, and period-halving bifurcation. Moreover, the effects of the impulsive perturbation and hatching rate on the chaos of the system are comparatively studied by numerical simulation. These obtained results can be useful for ecosystem management and for explaining complex phenomena of ecosystems.     

一类具有脉冲效应和HollingII型功能性反应的阶段结构的捕食系统

A stage-structured predator-prey system with impulsive effect and Holling type-II functional response is investigated. By the Floquet theory and small amplitude perturbation skills, it is proved that there exists a global stable pest-eradication periodic solution when the impulsive period is less than some critical values. Farther, the conditions for the permanence of system are established. Numerical simulations are carried out to illustrate the impulsive effect on the dynamics of the system.