The dynamical behaviors of a Ivlev-type two-prey two-predator system with impulsive effect

In this paper, considering the strategy of integrated Pest Management (IPM), a class of two-prey two-predator system with the Ivlev-type functional response and impulsive effect at different fixed time is established. By using impulsive comparison theorem, Floquent theory and small amplitude perturbation skill, the sufficient conditions for the system to be extinct of prey and permanence are proved. Moreover, we give two sufficient conditions for the extinction of one of two prey and remaining three species are permanent. Numerical simulation shows that there exist complex dynamics for system, such as symmetry-breaking pitchfork bifurcation, periodic doubling bifurcation, chaos, periodic halving cascade. Lastly, a brief discussion is given.     

Extinction and permanence of the predator-prey system with general functional response and impulsive control

Traditional approach for modelling the evolution of populations in the predator-prey ecosystem has commonly been undertaken using specific impulsive response function, and this kind of modelling is applicable only for a specific ecosystem under certain environmental situations only. This paper attempts to fill the gap by modelling the predator-prey ecosystem using a ‘generalized’ impulsive response function for the first time. Different from previous research, the present work develops the modelling for an integrated pest management (IPM) especially when the stocking of predator (natural enemy) and the harvesting of prey (pest) occur impulsively and at different instances of time. The paper firstly establishes the sufficient conditions for the local and the global stabilities of prey eradication periodic solution by applying the Floquet theorem of the Impulsive different equation and small amplitude perturbation under a ‘generalized’ impulsive response function. Subsequently the sufficient condition for the permanence of the system is given through the comparison techniques. The corollaries of the theorems that are established by using the ‘general impulsive response function’ under the locally asymptotically stable condition are found to be in excellent agreement with those reported previously. Theoretical results that are obtained in this work is then validated by using a typical impulsive response function (Holling type-II) as an example, and the outcome is shown to be consistent with the previously reported results. Finally, the implication of the developed theories for practical pest management is illustrated through numerical simulation. It is shown that the elimination of either the preys or the pest can be effectively deployed by making use of the theoretical model established in this work. The developed model is capable to predict the population evolutions of the predator-prey ecosystem to accommodate requirements such as: the combinations of the biological control, chemical control, any functional response function, the moderate impulsive period, the harvest rate for the prey and predator parameter and the incremental stocking of the predator parameter.     

Extinction and permanence in delayed stage-structure predator–prey system with impulsive effects

Based on the classical stage-structured model and Lotka–Volterra predator–prey model, an impulsive delayed differential equation to model the process of periodically releasing natural enemies at fixed times for pest control is proposed and investigated. We show that the conditions for global attractivity of the ‘pest-extinction’ (‘prey-eradication’) periodic solution and permanence of the population of the model depend on time delay. We also show that constant maturation time delay and impulsive releasing for the predator can bring great effects on the dynamics of system by numerical analysis. As a result, the pest maturation time delay is considered to establish a procedure to maintain the pests at an acceptably low level in the long term. In this paper, the main feature is that we introduce time delay and pulse into the predator–prey (natural enemy-pest) model with age structure, exhibit a new modelling method which is applied to investigate impulsive delay differential equations, and give some reasonable suggestions for pest management.     

Extinction and permanence of the predator–prey system with stocking of prey and harvesting of predator impulsively

In this paper, a predator–prey system with stocking of prey and harvesting of predator impulsively is studied. Here, the prey population is stocked with a constant quantity and the predator population is harvested at a rate proportional to the species itself at fixed moments. Under some conditions, the existence and global asymptotic stability of the boundary periodic solution are proved, which implies that the system will be extinct; and given some different restrictions, ultimate positive upper and lower bounds of all solutions are obtained, showing the system being permanent. At last, two examples are given to illustrate our results. Copyright © 2005 John Wiley & Sons, Ltd.     

Extinction and permanence in a stochastic non-autonomous population system

A stochastic non-autonomous predator–prey system with Holling II functional response is investigated. Sufficient criteria for extinction and uniform weak persistence in the mean for each species are established. The acute persistence–extinction thresholds for each species are obtained in many cases.     

Dynamics of a two-prey one-predator system with Watt-type functional response and impulsive control strategy

In this paper, by using theories and methods of ecology and ODE, a two-prey one-predator system with Watt-type functional response and impulsive perturbations on the predator is established. The system is affected by impulse which can be considered as a control. Conditions for the permanence of the system are obtained. The numerical analysis is carried out to study the effects of perturbation varying parameters of the system. The system shows the rich dynamic behavior including quasi-periodic oscillation, narrow periodic window, wide periodic window, chaotic bands, period doubling bifurcation, symmetry-breaking pitchfork bifurcation, period-halving bifurcation and crises, etc.     

Extinction and permanence in nonautonomous competitive system with stage structure

In this paper, the n-species nonautonomous stage-structured competitive system is constructed and considered. Sufficient conditions for its extinction and permanence are obtained. Results here generalize and unify some previous ones. Moreover, it is concluded that stage structure in this system is one of the important factors that effect the extinction and permanence of species.     

The effect of dispersal on the permanence of a predator–prey system with time delay

A predator–prey model with prey dispersal and time delay due to the gestation of the predator is investigated. By analyzing the corresponding characteristic equations, the local stability of each of the nonnegative equilibria is discussed. By using an iteration technique, a threshold is derived for the permanence and extinction of the proposed model. Numerical simulations are carried out to illustrate the main results.     

基于脉冲扰动作用下一个捕食者-两个食饵模型的动力学性质

基于害虫的生物控制策略,分别利用Floquet乘子理论及脉冲比较定理,研究了一类具有脉冲效应的一个捕食者-两个食饵模型并进行了分析,得到害虫根除周期解的渐近稳定与系统持续生存条件.     

Extinction and stability of an impulsive system with pure delays

In this paper, we consider an impulsive competitive system with infinite delays and investigate the extinction and stability of the system. For the corresponding impulsive logistic model, our stability conditions are weaker than those of Yang et al. (2011).     

The study of predator–prey system with defensive ability of prey and impulsive perturbations on the predator

Predator–prey system with non-monotonic functional response and impulsive perturbations on the predator is established. By using Floquet theorem and small amplitude perturbation skills, a locally asymptotically stable prey-eradication periodic solution is obtained when the impulsive period is less than the critical value. Otherwise, if the impulsive period is larger than the critical value, the system is permanent. Further, using numerical simulation method the influences of the impulsive perturbations on the inherent oscillation are investigated. With the increasing of the impulsive value, the system displays a series of complex phenomena, which include (1) quasi-periodic oscillating, (2) period-doubling, (3) period-halfing, (4) non-unique dynamics (meaning that several attractors coexist), (5) attractor crisis and (6) chaotic bands with periodic windows.     

Dynamics of a periodic Watt-type predator–prey system with impulsive effect

In this paper, an impulsive periodic predator–prey system with Watt-type functional response is investigated. By using the Floquet theory of linear periodic impulsive equation, the stability conditions for the prey-eradication positive periodic solution are given, and the boundedness of the system is proved. By the method of coincidence degree, the sufficient conditions for the existence of at least one strictly positive periodic solution are obtained. Furthermore, we give numerical analysis to confirm our theoretical results. It will be useful for ecosystem control.     

Extinction and permanence of chemostat model with pulsed input in a polluted environment

In this paper, chemostat model with pulsed input in a polluted environment is considered. By using the Floquet theorem, we find the microorganism eradication periodic solution is globally asymptotically stable if some conditions are needed. At the same time we can find the condition of the nutrient and microorganism are permanent.     

具有避难所的非自治两种群捕食者-食饵系统捕食者种群绝灭性研究

研究一类食饵具有避难所的非自治两种群捕食者-食饵系统,借助微分方程振荡性理论和微分方程比较原理得到了保证捕食者绝灭的一组充分性条件.     

Extinction and permanence for a pulse vaccination delayed SEIRS epidemic model

A delayed SEIRS epidemic model with pulse vaccination and bilinear incidence rate is investigated. Using Krasnoselskii’s fixed-point theorem, we obtain the existence of disease-free periodic solution (DFPS for short) of the delayed impulsive epidemic system. Further, using the comparison method, we prove that under the condition R^* < 1, the DFPS is globally attractive, and that R_* > 1 implies that the disease is permanent. Theoretical results show that the disease will be extinct if the vaccination rate is larger than θ^* and the disease is uniformly persistent if the vaccination rate is less than θ_*. Our results indicate that a long latent period of the disease or a large pulse vaccination rate will lead to eradication of the disease.     

Global attractability and permanence for a new stage-structured delay impulsive ecosystem

In this paper, a new stage-structured delay ecosystem with impulsive effect is formulated and some dynamical properties of this system is investigated.By using comparison theorem and the stroboscopic technique, we prove the existence of the predator-extinction periodic solution of this system and obtain some sufficient conditions to guarantee the global attractivity of the prey-extinction periodic solution. In the final, we also obtain the permanence of this system. It should be pointed out that the new mathematical method used in this paper can also be applied to investigate such other ecosystems corresponding to both impulsive and delay differential equations.     

Dynamics of an impulsive control system which prey species share a common predator

In an ecosystem multiple prey species often share a common predator and the interactions between the preys are neutral. In view of these facts and based on a multiple species prey–predator system with Holling IV and II functional responses, an impulsive differential equation to model the process of periodically releasing natural enemies and spraying pesticides at different fixed times for pest control is proposed and investigated. It is proved that there exists a locally asymptotically stable pest-eradication periodic solution under the assumption that the impulsive period is less than some critical value (or the release amount of the predator is greater than another critical value). Permanence conditions are established when the impulsive period is greater than another critical value (or the release amount of the predator is less than some critical value). Numerical results show that the system we consider has more complex dynamics including period solution, quasi-periodic oscillation, chaos, intermittency and crises.     

Weak average persistence and extinction of a predator–prey system in a polluted environment with impulsive toxicant input

In this paper, we have investigated a predator–prey system in a polluted environment with impulsive toxicant input at fixed moments. We have obtained two thresholds on the impulsive period by assuming the toxicant amount input is fixed to the environment at each pulse moment. If the impulsive period is greater than the big threshold, then both populations are weak average persistent. If the period lies between of the two thresholds, then the prey population will be weak average persistent while the predator population extinct. If the period is less than the small threshold, both populations tend to extinction. Finally, our theoretical results are confirmed by own numerical simulations.