Frequency discrimination for pulsed versus modulated tones

Estimates of frequency discrimination for pulsed modulated tones were obtained by 11 observers at 350, 500, 1000, 4000, and 8000 Hz. At low frequencies, frequency DL's are larger for modulated than for pulsed tones; at 8000 Hz the contrary was found. Frequency DL's (difference limens) determined by different methods and procedures differed by a factor up to four; extreme individual frequency DL's, however, by a factor up to 27.     

Intensity discrimination of Gaussian-windowed tones: indications for the shape of the auditory frequency-time window.

The just-noticeable difference in intensity jnd(I) was measured for 1-kHz tones with a Gaussian-shaped envelope as a function of their spectro-temporal shape. The stimuli, with constant energy and a constant product of bandwidth and duration, ranged from a long-duration narrow-band "tone" to a short-duration broadband "click." The jnd(I) was measured in three normal-hearing listeners at sensation levels of 0, 10, 20, and 30 dB in 35 dB(A) SPL pink noise. At intermediate sensation levels, jnd(I) depends on the spectro-temporal shape: at the extreme shapes (tones and clicks), intensity discrimination performance is best, whereas at intermediate shapes the jnd(I) is larger. Similar results are observed at a higher overall sound level, and at a higher carrier frequency. The maximum jnd(I) is observed for stimuli with an effective bandwidth of about 1/3 octave and an effective duration of 4 ms at 1 kHz (1 ms at 4 kHz). A generalized multiple-window model is proposed that assumes that the spectro-temporal domain is partitioned into "internal" auditory frequency-time windows. The model predicts that intensity discrimination thresholds depend upon the number of windows excited by a signal: jnd(I) is largest for stimuli covering one window.     

Detection and F0 discrimination of harmonic complex tones in the presence of competing tones or noise

Normal-hearing listeners' ability to "hear out" the pitch of a target harmonic complex tone (HCT) was tested with simultaneous HCT or noise maskers, all bandpass-filtered into the same spectral region (1200-3600 Hz). Target-to-masker ratios (TMRs) necessary to discriminate fixed fundamental-frequency (F0) differences were measured for target F0s between 100 and 400 Hz. At high F0s (400 Hz), asynchronous gating of masker and signal, presenting the masker in a different F0 range, and reducing the F0 rove of the masker, all resulted in improved performance. At the low F0s (100 Hz), none of these manipulations improved performance significantly. The findings are generally consistent with the idea that the ability to segregate sounds based on cues such as F0 differences and onset/offset asynchronies can be strongly limited by peripheral harmonic resolvability. However, some cases were observed where perceptual segregation appeared possible, even when no peripherally resolved harmonics were present in the mixture of target and masker. A final experiment, comparing TMRs necessary for detection and F0 discrimination, showed that F0 discrimination of the target was possible with noise maskers at only a few decibels above detection threshold, whereas similar performance with HCT maskers was only possible 15-25 dB above detection threshold.     

Frequency discrimination of complex tones with overlapping and non-overlapping harmonics

These experiments address the following issues. (1) When two complex tones contain different harmonics, do the differences in timbre between them impair the ability to discriminate the pitches of the tones? (2) When two complex tones have only a single component in common, and that component is the most discriminable component in each tone, is the frequency discrimination of the component affected by differences in residue pitch between the two tones? (3) How good is the pitch discrimination of complex tones with no common components when each tone contains multiple harmonics, so as to avoid ambiguity of pitch? (4) Is the pitch discrimination of complex tones with common harmonics impaired by shifting the component frequencies to nonharmonic values? In all experiments, frequency difference limens (DLCs) were measured for multiple-component complex tones, using an adaptive two-interval, two-alternative, forced-choice task. Three highly trained subjects were used. The main conclusions are as follows. (1) When two tones have the first six harmonics in common, DLCs are larger when the upper harmonics are different than when the upper harmonics are in common or are absent. It appears that differences in timbre impair DLCs. (2) Discrimination of the frequency of a single common partial in two complex tones is worse when the two tones have different residue pitches than when they have the same residue pitch. (3) DLCs for complex tones with no common harmonics are generally larger than those for complex tones with common harmonics. For the former, large individual differences occur, probably because subjects are affected differently by differences in timbre. (4) DLCs for harmonic complex tones are smaller than DLCs for complex tones in which the components are mistuned from harmonic values. This can probably be attributed to the less distinct residue pitch of the inharmonic complexes, rather than to reduced discriminability of partials. Overall, the results support the idea that DLCs for complex tones with common harmonics depend on residue pitch comparisons, rather than on comparisons of the pitches of partials.     

Loudness of brief tones measured by magnitude estimation and loudness matching

McFadden [J. Acoust. Soc. Am. 57, 702-704 (1975)] questioned the accuracy and reliability of magnitude estimation for measuring loudness of tones that vary both in duration and level, whereas Stevens and Hall [Percept. Psychophys. 1, 319-327 (1966)] reported reasonable group data. To gain insight into this discrepancy, the present study compares loudness measures for 5- and 200-ms tones using magnitude estimation and equal-loudness matches from the same listeners. Results indicate that both procedures provide rapid and accurate assessments of group loudness functions for brief tones, but may not be reliable enough to reveal specific characteristics of loudness in individual listeners.     

Frequency discrimination of tones presented in filtered noise

Previous research (Emmerich et al., 1983) in which tones were presented in the center of the notches in band-reject noise backgrounds suggests that information from frequency regions remote from the nominal signal frequency is useful in frequency discrimination. The present work extends the earlier findings by presenting tones on either side of a notch so that only one (or the other) tail of the excitation patterns of the tones would fall into the notch. In addition, tones were presented in high-pass noise, low-pass noise, and various combinations of the two. The results again indicate that remote information affects frequency discrimination, and they are also consistent with the hypothesis that the low-frequency tail of the excitation pattern is more useful for frequency discrimination than is the high-frequency tail.     

Neural mechanisms of interstimulus interval-dependent responses in the primary auditory cortex of awake cats

Background  

Primary auditory cortex (AI) neurons show qualitatively distinct response features to successive acoustic signals depending on the inter-stimulus intervals (ISI). Such ISI-dependent AI responses are believed to underlie, at least partially, categorical perception of click trains (elemental vs. fused quality) and stop consonant-vowel syllables (eg.,/da/-/ta/continuum).     

Intensity discrimination of pulsed tones by the goldfish (Carassius auratus)

Intensity discrimination thresholds for 500-ms pure-tone bursts were measured as a function of frequency in the goldfish (Carassius auratus) using classical respiratory conditioning. At 55-dB sensation level (SL), thresholds range from 1.44-2.2 dB between 100 and 1600 Hz. There is not important effect of frequency on intensity discrimination. Thresholds at 35-dB SL average 0.7 dB higher than at 55-dB SL. This is a small difference in the context of the threshold variability. In intensity discrimination acuity, the goldfish is quantitatively similar to other vertebrates, including birds and mammals.     

Responses to amplitude-modulated tones in the auditory nerve of the cat.

Sinusoidally amplitude-modulated (AM) tones are frequently used in psychophysical and physiological studies, yet a comprehensive study on the coding of AM tones in the auditory nerve is lacking. AM responses of single auditory-nerve fibers of the cat are studied, systematically varying modulation depth, frequency, and sound level. Synchrony-level functions were nonmonotonic with maximum values that were inversely correlated with spontaneous rate (SR). In most fibers, envelope phase-locking showed a positive gain. Modulation transfer functions were uniformly low pass. Their corner frequency increased with characteristic frequency (CF), but changed little for CFs above 10 kHz. The highest modulation frequencies to which phase locking occurred were more than 0.8 oct lower than the highest frequencies to which phase locking to pure tones occurs. Cumulative, or unwrapped, phase increased linearly with modulation frequency: The slope was inversely related to CF, and slightly higher than group delays reported for pure tones. High SR, low CF fibers showed the poorest envelope phase locking. In some low CF fibers, phase locking increased at high levels, associated with "peak-splitting" phenomena. Changes in average rate due to modulation were small, and could be enhancement or suppression.     

Reliability of pitch discrimination of vowel and piano tones

This study evaluated the reliability of pitch judgments as a basic step toward increasing interrater and intrarater reliability of multidimensional perceptual judgments of the speaking voice. Forty-five undergraduate university students studying speech/language pathology made piano-to-piano tone pitch matches and vowel-to-piano pitch matches using a computer software program. The mean percentage correct of piano-to-piano tone matches was 91.3% and of vowel-to-piano matches was 75.6%. Subjects who scored 100% correct were significantly faster at the pitch matching task. Further research of perceptual judgments of pitch and its contribution to multidimensional rating tasks is warranted.     

Level discrimination of tones as a function of duration

Difference limens for level [delta Ls (dB) = 20 log[p + delta p)/p), where p is the pressure] were measured as a function of duration for tones at 250, 500, and 8000 Hz. Stimulus durations ranged from 2 ms to 2 s, and the stimulus power was held constant. Rise and fall times were 1 ms. The interstimulus interval was 250 ms. At each frequency, three levels were tested: 85, 65, and approximately 40 dB SPL. An adaptive two-alternative forced-choice procedure with feedback was used. For three normal listeners, delta Ls decreased as duration increased, up to at least 2 s, except at 250 Hz. At 250 Hz, delta L stopped decreasing at durations between 0.5 and 1 s. In a double logarithmic plot of delta L versus duration, the rate of decrease is generally well fitted by a sloping line. The average slope is -0.28; it is steeper at high levels than at low levels. Because the average slope is shallower than the -0.5 slope predicted for an optimum detector, it may be that fast adaptation of auditory-nerve activity and/or memory effects interfere with level discrimination of long-duration tones. Finally, the delta Ls at 8 kHz decreased nonmonotonically with increasing level.     

Detection and discrimination of simulated motion of auditory targets in the horizontal plane

Three experiments investigated subjects' ability to detect and discriminate the simulated horizontal motion of auditory targets in an anechoic environment. "Moving" stimuli were produced by dynamic application of stereophonic balancing algorithms to a two-loudspeaker system with a 30 degree separation. All stimuli were 500-Hz tones. In experiment 1, subjects had to discriminate a left-to-right moving stimulus from a stationary stimulus pulsed for the same duration (300 or 600 ms). For both durations, minimum audible "movement" angles ("MAMA's") were on the order of 5 degrees for stimuli presented at 0 degrees azimuth (straight ahead), and increased to greater than 30 degrees for stimuli presented at +/- 90 degrees azimuth. Experiment 2 further investigated MAMA's at 0 degrees azimuth, employing two different procedures to track threshold: holding stimulus duration constant (at 100-600 ms) while varying velocity; or holding the velocity constant (at 22 degrees-360 degrees/s) while varying duration. Results from the two procedures agreed with each other and with the MAMA's determined by Perrott and Musicant for actually moving sound sources [J. Acoust. Soc. Am. 62, 1463-1466 (1977b)]: As stimulus duration decreased below 100-150 ms, the MAMA's increased sharply from 5 degrees-20 degrees or more, indicating that there is some minimum integration time required for subjects to perform optimally in an auditory spatial resolution task. Experiment 3 determined differential "velocity" thresholds employing simulated reference velocities of 0 degrees-150 degrees/s and stimulus durations of 150-600 ms. As with experiments 1 and 2, the data are more easily summarized by considering angular distance than velocity: For a given "extent of movement" of a reference target, about 4 degrees-10 degrees additional extent is required for threshold discrimination between two "moving" targets, more or less independently of stimulus duration or reference velocity. These data suggest that for the range of simulated velocities employed in these experiments, subjects respond to spatial changes--not velocity per se--when presented with a "motion" detection or discrimination task.     

Aging and temporal discrimination in auditory sequences.

This study examined age-related changes in temporal sensitivity to increments in the inter-onset intervals (IOI) of successive components in tonal sequences. Temporal discrimination was examined using reference stimulus patterns consisting of five 50-ms, 4000-Hz components with equal tonal IOIs selected from the range 100-600 ms. Discrimination was examined in separate conditions by measuring the relative difference limen (DL) for increments of tonal IOI in comparison sequences. In some conditions, comparison sequences featured equal increments of all tonal lOIs to examined listener sensitivity to uniform changes of sequence rate, or tempo. Other conditions measured the DL for increments of a single target IOI within otherwise uniform-rate comparison sequences. For these measurements, the single target IOI was either fixed in sequence location, or randomized in location across listening trials. Listeners in the study included four groups of young and elderly adults with and without high-frequency hearing loss. The results for all listeners showed the relative DL for rate discrimination to decrease from a maximum at the 100-ms IOI to a smaller stable value across the range of longer sequence IOI. All listeners also exhibited larger relative DLs for discrimination of single target intervals compared to rate discrimination for equivalent reference IOI values. Older listeners showed poorer performance than younger listeners in all conditions, with the largest age differences observed for discrimination of brief single intervals that were varied randomly in sequence location. None of the results revealed significant effects of hearing loss on performance of younger and older listeners.     

Level discrimination of single tones in a multitone complex by normal-hearing and hearing-impaired listeners

A conditional-on-a-single-stimulus (COSS) analysis procedure [B. G. Berg, J. Acoust. Soc. Am. 86, 1743-1746 (1989)] was used to estimate how well normal-hearing and hearing-impaired listeners selectively attend to individual spectral components of a broadband signal in a level discrimination task. On each trial, two multitone complexes consisting of six octave frequencies from 250 to 8000 Hz were presented to listeners. The levels of the individual tones were chosen independently and at random on each presentation. The target tone was selected, within a block of trials, as the 250-, 1000-, or 4000-Hz component. On each trial, listeners were asked to indicate which of the two complex sounds contained the higher level target. As a group, normal-hearing listeners exhibited greater selectivity than hearing-impaired listeners to the 250-Hz target, while hearing-impaired listeners showed greater selectivity than normal-hearing listeners to the 4000-Hz target, which is in the region of their hearing loss. Both groups of listeners displayed large variability in their ability to selectively weight the 1000-Hz target. Trial-by-trial analysis showed a decrease in weighting efficiency with increasing frequency for normal-hearing listeners, but a relatively constant weighting efficiency across frequency for hearing-impaired listeners. Interestingly, hearing-impaired listeners selectively weighted the 4000-Hz target, which was in the region of their hearing loss, more efficiently than did the normal-hearing listeners.     

Detection of tones in reproducible narrow-band noise

Hit and false-alarm rates were measured for detection of a 500-Hz tone target in each of ten reproducible samples of 1/3-oct bandwidth noise centered at 500 Hz for both NoS pi and NoSo conditions. The effects on hit rates of the starting phase of the target relative to individual noise samples were investigated with two target phase angles for three subjects. The major results are: (1) performance varies significantly over masker waveforms; (2) for NoS pi conditions, the effect of target-to-marker phase angle on hit rates is not significant for these narrow-band maskers; (3) for NoSo conditions, the target-to-masker phase angle has a large effect; (4) no significant correlation between NoSo performance and NoS pi performance is seen across masker waveforms. These results are generally consistent wuth previously reported results for wideband maskers [R.H. Gilkey, D.E. Robinson, and T.E. Hanna, "Effects of masker waveform and signal-to-masker phase relation on diotic and dichotic masking by reproducible noise," J. Acoust. Soc. Am. 78, 1207-1219 (1985)] with an important exception. Specifically, in the wideband experiment, significant correlation between NoSo and NoS pi performance across noise samples was found. In addition, in the wideband experiment, a small yet statistically significant effect of target-to-masker phase was observed in the NoS pi condition.     

The functional analysis of auditory discrimination

Mammals have evolved the ability to acquire auditory discriminations. The characteristics of this discriminative ability presumably fit the natural conditions under which discriminations are normally acquired. The purpose of this paper is to review experiments which were directed at showing that auditory discriminations are most rapidly acquired when natural features are incorporated into the experiments. The experiments were also directed at discovering the underlying characteristics of the discriminative ability. When animals were trained to discriminate the position of a sound source in which natural features were incorporated into the experiment, the discrimination was acquired in one trial. Manipulation of the natural features suggested that one trial acquisition depends upon the following. (1) Stimulus novelty; the effect of reinforcement is stronger in the presence of novel than familiar stimuli. (2) Specific behavioral effect of reinforcement; the effect of reinforcing a response in the presence of a novel auditory stimulus is to increase the strength of approaching and manipulating the sound source.     

Complex tone processing in primary auditory cortex of the awake monkey. II. Pitch versus critical band representation

Noninvasive neurophysiological studies in humans support the existence of an orthogonal spatial representation of pure tone frequency and complex tone pitch in auditory cortex [Langner et al., J. Comp. Physiol. A 181, 665-676 (1997)]. However, since this topographic organization is based on neuromagnetic responses evoked by wideband harmonic complexes (HCs) of variable fundamental frequency (f0), and thus interharmonic frequency separation (deltaF), critical band filtering effects due to differential resolvability of harmonics may have contributed to shaping these responses. To test this hypothesis, the present study examined responses evoked by three-component HCs of variable f0 in primary auditory cortex (A1) of the awake monkey. The center frequency of the HCs was fixed at the best frequency (BF) of the cortical site. Auditory evoked potential (AEP), multiunit activity, and current source density techniques were used to evaluate A1 responses as a function of f0 (=deltaF). Generally, amplitudes of nearly all response components increased with f0, such that maximal responses were evoked by HCs comprised of low-order resolved harmonics. Statistically significant increases in response amplitude typically occurred at deltaFs between 10% and 20% of center frequency, suggestive of critical bandlike behavior. Complex tone response amplitudes also reflected nonlinear summation in that they could not be predicted by the pure tone frequency sensitivity curves of the cortical sites. A mechanism accounting for the observed results is proposed which involves mutual lateral inhibitory interactions between responses evoked by stimulus components lying within the same critical band. As intracortical AEP components likely to be propagated to the scalp were also strongly modulated by deltaF, these findings indicate that noninvasive recordings of responses to complex sounds may require a consideration of critical band effects in their interpretation.     

Repetition rate and signal level effects on neuronal responses to brief tone pulses in cat auditory cortex

This study describes the effects on the spike count, spike timing, and entrainment of cat auditory cortex neurons of parametric variations in the repetition rate and amplitude of a brief, characteristic frequency tone pulse. Data were obtained from single neurons in barbiturate-anesthetized cats to which signals were presented monaurally to the ear contralateral to the recording electrode. All neurons showed low-pass sensitivity to tone repetition rate. In cells with a monotonic rate response, the effect of an increasing stimulus level was to elevate the response rate and to extend performance to higher repetition rates. In nonmonotonic cells, cutoff frequencies (for repetition rate) varied with overall spike count. Latent periods increased with increases in repetition rate. This effect developed over the first few stimulus trials at any given repetition rate. Spike entrainment to the tone pulses varied with both repetition rate and signal level. Increases in signal level improved entrainment for responses to stimuli presented at low repetition rates, but entrainment at high repetition rates always saturated at significantly imperfect levels.     

A high-precision magnetoencephalographic study of human auditory steady-state responses to amplitude-modulated tones

The cerebral magnetic field of the auditory steady-state response (SSR) to sinusoidal amplitude-modulated (SAM) tones was recorded in healthy humans. The waveforms of underlying cortical source activity were calculated at multiples of the modulation frequency using the method of source space projection, which improved the signal-to-noise ratio (SNR) by a factor of 2 to 4. Since the complex amplitudes of the cortical source activity were independent of the sensor position in relation to the subject's head, a comparison of the results across experimental sessions was possible. The effect of modulation frequency on the amplitude and phase of the SSR was investigated at 30 different values between 10 and 98 Hz. At modulation frequencies between 10 and 20 Hz the SNR of harmonics near 40 Hz were predominant over the fundamental SSR. Above 30 Hz the SSR showed an almost sinusoidal waveform with an amplitude maximum at 40 Hz. The amplitude decreased with increasing modulation frequency but was significantly different from the magnetoencephalographic (MEG) background activity up to 98 Hz. Phase response at the fundamental and first harmonic decreased monotonically with increasing modulation frequency. The group delay (apparent latency) showed peaks of 72 ms at 20 Hz, 48 ms at 40 Hz, and 26 ms at 80 Hz. The effects of stimulus intensity, modulation depth, and carrier frequency on amplitude and phase of the SSR were also investigated. The SSR amplitude decreased linearly when stimulus intensity or the modulation depth were decreased in logarithmic steps. SSR amplitude decreased by a factor of 3 when carrier frequency increased from 250 to 4000 Hz. From the phase characteristics, time delays were found in the range of 0 to 6 ms for stimulus intensity, modulation depth, and carrier frequency, which were maximal at low frequencies, low intensities, or maximal modulation depth.