Extinction and permanence in delayed stage-structure predator–prey system with impulsive effects

Based on the classical stage-structured model and Lotka–Volterra predator–prey model, an impulsive delayed differential equation to model the process of periodically releasing natural enemies at fixed times for pest control is proposed and investigated. We show that the conditions for global attractivity of the ‘pest-extinction’ (‘prey-eradication’) periodic solution and permanence of the population of the model depend on time delay. We also show that constant maturation time delay and impulsive releasing for the predator can bring great effects on the dynamics of system by numerical analysis. As a result, the pest maturation time delay is considered to establish a procedure to maintain the pests at an acceptably low level in the long term. In this paper, the main feature is that we introduce time delay and pulse into the predator–prey (natural enemy-pest) model with age structure, exhibit a new modelling method which is applied to investigate impulsive delay differential equations, and give some reasonable suggestions for pest management.     

The dynamics of pest control pollution model with age structure and time delay

In this paper, by using pollution model and impulsive delay differential equation, we investigate the dynamics of a pest control model with age structure for pest by introducing a constant periodic pesticide input and releasing natural enemies at different fixed moment. We assume only the pests are affected by pesticide. We show that there exists a global attractive pest-extinction periodic solution when the periodic natural enemies release amount μ₁ and pesticide input amount μ₂ are larger than some critical value. Further, the condition for the permanence of the system is also given. By numerical analyses, we also show that constant maturation time delay, pulse pesticide input and pulse releasing of the natural enemies can bring obvious effects on the dynamics of system. We believe that the results will provide reliable tactic basis for the practical pest management.     

Dynamics of an impulsive control system which prey species share a common predator

In an ecosystem multiple prey species often share a common predator and the interactions between the preys are neutral. In view of these facts and based on a multiple species prey–predator system with Holling IV and II functional responses, an impulsive differential equation to model the process of periodically releasing natural enemies and spraying pesticides at different fixed times for pest control is proposed and investigated. It is proved that there exists a locally asymptotically stable pest-eradication periodic solution under the assumption that the impulsive period is less than some critical value (or the release amount of the predator is greater than another critical value). Permanence conditions are established when the impulsive period is greater than another critical value (or the release amount of the predator is less than some critical value). Numerical results show that the system we consider has more complex dynamics including period solution, quasi-periodic oscillation, chaos, intermittency and crises.     

Eco-epidemiology model with age structure and prey-dependent consumption for pest management

In this paper, a prey-dependent consumption predator–prey (natural enemy-pest) model with age structure for the predators and infectious disease in the prey, is considered. Infectious pests, immature natural enemies and mature natural enemies are released impulsively. By using Floquet’s theorem, small-amplitude perturbation skills and comparison theorem, we obtain both the sufficient conditions for the global asymptotical stability of the susceptible pest-eradication periodic solution and the permanence of the system. The results provide a reliable theoretical tactics for pest management.     

Impulsive control strategy of a pest management SI model with nonlinear incidence rate

In this work, we consider a pest management SI model with impulsive release of infective pests and spraying pesticides. We prove that all solutions of the investigated system are uniformly ultimately bounded and the pest-extinction periodic solution is globally asymptotically stable when some condition is satisfied. We also obtain the permanent condition of the system. It is concluded that the approach of combining impulsive release of infective pests with impulsive spraying pesticides provides reliable tactic basis for the practical pest management.     

Hybrid approach for pest control with impulsive releasing of natural enemies and chemical pesticides: A plant–pest–natural enemy model

The agricultural pests can be controlled effectively by simultaneous use (i.e., hybrid approach) of biological and chemical control methods. Also, many insect natural enemies have two major life stages, immature and mature. According to this biological background, in this paper, we propose a three tropic level plant–pest–natural enemy food chain model with stage structure in natural enemy. Moreover, impulsive releasing of natural enemies and harvesting of pests are also considered. We obtain that the system has two types of periodic solutions: plant–pest-extinction and pest-extinction using stroboscopic maps. The local stability for both periodic solutions is studied using the Floquet theory of the impulsive equation and small amplitude perturbation techniques. The sufficient conditions for the global attractivity of a pest-extinction periodic solution are determined by the comparison technique of impulsive differential equations. We analyze that the global attractivity of a pest-extinction periodic solution and permanence of the system are evidenced by a threshold limit of an impulsive period depending on pulse releasing and harvesting amounts. Finally, numerical simulations are given in support of validation of the theoretical findings.     

Dynamics on a pest management SI model with control strategies of different frequencies

Based on spraying pesticide and introducing infected pest and natural enemy for pest control, an SI ecological epidemic model with different frequencies of pesticide applications and infected pests and natural enemy releases is proposed and studied. With spraying either more or less frequently than the releases, the threshold condition of existence and global attractiveness of susceptible pest extinction periodic solution is obtained. We investigate the effects of the pest control tactics on the threshold conditions. We also show that the system has rich dynamics including period-doubling bifurcations and chaos as the release period increases, which implies that the presence of impulsive intervention makes the dynamic behavior more complex. Finally, to see how the pesticide applications can be reduced, we develop a model involving periodic releases of natural enemies with chemical control applied only when the densities of the pest reaches the given Economic Threshold. It indicates that the hybrid method is the most effective method to control pest and the frequency of pesticide applications largely depends on the initial densities and the control tactics.     

具有脉冲效应和综合害虫控制的捕食系统

本文通过生物控制和化学控制提出了具有周期脉冲效应与害虫控制的捕食系统. 系统保护天敌避免灭绝,在一些条件下可以使害虫灭绝.就是说当脉冲周期小于某一临界值时,存在全局稳定害虫灭绝周期解.脉冲周期增大大于临界值时,平凡害虫灭绝周期解失去稳定性并产生正周期解,利用分支理论来研究正周期解的存在性.进而,利用李雅普诺夫函数和比较定理确定了持续生存的条件.     

Dynamic analysis of pest control model with population dispersal in two patches and impulsive effect

In this paper, we investigate the pest control model with population dispersal in two patches and impulsive effect. By exploiting the Floquet theory of impulsive differential equation and small amplitude perturbation skills, we can obtain that the susceptible pest eradication periodic solution is globally asymptotically stable if the impulsive periodic τ is less than the critical value τ₀ . Further, we also prove that the system is permanent when the impulsive periodic τ is larger than the critical value τ₀. Hence, in order to drive the susceptible pest to extinction, we can take impulsive control strategy such that τ < τ₀ according to the effect of the viruses on the environment and the cost of the releasing pest infected in a laboratory. Finally, numerical simulations validate the obtained theoretical results for the pest control model with population dispersal in two patches and impulsive effect.     

Models for determining how many natural enemies to release inoculatively in combinations of biological and chemical control with pesticide resistance

Combining biological and chemical control has been an efficient strategy to combat the evolution of pesticide resistance. Continuous releases of natural enemies could reduce the impact of a pesticide on them and the number to be released should be adapted to the development of pesticide resistance. To provide some insights towards this adaptation strategy, we developed a novel pest–natural enemy model considering both resistance development and inoculative releases of natural enemies. Three releasing functions which ensure the extinction of the pest population are proposed and their corresponding threshold conditions obtained. Aiming to eradicate the pest population, an analytic formula for the number of natural enemies to be released was obtained for each of the three different releasing functions, with emphasis on their biological implications. The results can assist in the design of appropriate control strategies and decision-making in pest management.     

A delayed epidemic model with stage-structure and pulses for pest management strategy

From a biological pest management standpoint, epidemic diseases models have become important tools in control of pest populations. This paper deals with an impulsive delay epidemic disease model with stage-structure and a general form of the incidence rate concerning pest control strategy, in which the pest population is subdivided into three subgroups: pest eggs, susceptible pests, infectious pests that do not attack crops. Using the discrete dynamical system determined by the stroboscopic map, we obtain the exact periodic susceptible pest-eradication solution of the system and observe that the susceptible pest-eradication periodic solution is globally attractive, provided that the amount of infective pests released periodically is larger than some critical value. When the amount of infective pests released is less than another critical value, the system is shown to be permanent, which implies that the trivial susceptible pest-eradication solution loses its attractivity. Our results indicate that besides the release amount of infective pests, the incidence rate, time delay and impulsive period can have great effects on the dynamics of our system.     

Mathematical modeling and control of population systems: Applications in biological pest control

The aim of this paper is to apply methods from optimal control theory, and from the theory of dynamic systems to the mathematical modeling of biological pest control. The linear feedback control problem for nonlinear systems has been formulated in order to obtain the optimal pest control strategy only through the introduction of natural enemies. Asymptotic stability of the closed-loop nonlinear Kolmogorov system is guaranteed by means of a Lyapunov function which can clearly be seen to be the solution of the Hamilton–Jacobi–Bellman equation, thus guaranteeing both stability and optimality. Numerical simulations for three possible scenarios of biological pest control based on the Lotka–Volterra models are provided to show the effectiveness of this method.     

A impulsive infective transmission SI model for pest control

In this paper, we propose a model with impulsive control of epidemics for pest management. By using Floquet's theorem, small‐amplitude perturbation skills and comparison theorem, we show that there exists a globally asymptotically stable susceptible pest‐eradication periodic solution when the release amount of infective pests is larger than some critical value. However, when the amount of infective pests released is less than this critical value, the system is shown to be permanent, which implies that the trivial periodic susceptible pest‐eradication solution loses its stability. Further, the existence of a positive periodic endemic solution and other rich dynamics are also studied by numerical simulation. Therefore, we can use the amount of release of infective pests to control susceptible pests at desirable low levels. Copyright © 2007 John Wiley & Sons, Ltd.     

害虫治理的病毒感染模型

研究了食饵受病毒感染且捕食者具有Beddington-DeAngelis功能性反应的生态流行病模型,此模型考虑的是脉冲释放病毒颗粒和自然天敌. 利用Floquet乘子理论、小振幅扰动技巧和比较定理证明了害虫根除周期解的全局渐近稳定性以及系统持续生存的充分条件.结论为现实的害虫管理提供了有效的策略依据.     

Impulsive control for a predator-prey Gompertz system with stage structure

For pest control in agriculture, we investigate the dynamics of a stage-structured predator-prey Gompertz system with impulsive spraying pesticide and releasing of natural enemies at different fixed moment. Using the stroboscopic map and comparison theorem, we obtain the sufficient conditions for the global attractivity of the mature predator-extinction periodic solution and the permanence of the system. Numerical simulations are inserted to verify the feasibility of the theoretical results, which show that the impulsive control plays a key role on the permanence of the system and also provide tactical basis for pest control.     

Dynamic analysis of a pest management SEI model with saturation incidence concerning impulsive control strategy

According to biological strategy for pest control, we investigate the dynamic behavior of a pest management SEI model with saturation incidence concerning impulsive control strategy-periodic releasing infected pests at fixed times. We prove that all solutions of the system are uniformly ultimately bounded and there exists a globally asymptotically stable pest-eradication periodic solution when the impulsive period is less than some critical value. When the impulsive period is larger than some critical value, the stability of the pest-eradication periodic solution is lost; the system is uniformly permanent. Thus, we can use the stability of the positive periodic solution and its period to control insect pests at acceptably low levels. Numerical results show that the system we consider can take on various kinds of periodic fluctuations and several types of attractor coexistence and is dominated by period-doubling cascade, symmetry-breaking pitchfork bifurcation, quasi-periodic oscillate, chaos, and non-unique dynamics.     

Optimal timing of interventions in fishery resource and pest management

Assuming that a fish population follows the continuous logistic growth or the discrete Beverton-Holt model, several optimal impulsive harvesting policies for the maximum stock level of the fish at the end of a fishing season are investigated under the condition of fixed intensity and frequency of impulsive harvesting. The optimal impulsive harvesting moments for all cases considered are given analytically and the related numerical simulations are also provided. Furthermore, the methods employed can also be used to investigate the optimal timing of chemical control in pest management. Our results confirm that the optimal timing of pesticide applications such that the density of the pest population is minimal at any time during a planting season or the average of density of the pest population over the planting season is minimal is the beginning of the planting season. In practice, the results can be used to guide the fisherman to manage fisheries and guide farmers to control pests.     

基于喷洒杀虫剂及释放病虫的脉冲控制害虫模型

基于喷洒杀虫剂及释放病虫的综合控制害虫策略,建立了具有脉冲控制的微分方程模型.利用脉冲微分方程的F loquet理论、比较定理,证明了害虫灭绝周期解的全局渐近稳定性与系统的持久性.     

Geometrical analysis of a pest management model in food-limited environments with nonlinear impulsive state feedback control

In this paper, a nonlinear impulsive state feedback control system is proposed to model an integrated pest management in food-limited environments. In the system, impulsive feedback control measures are implemented to control pests on the basis of the quantitative state of pests. Mathematically, an intuitive geometric analysis is used to indicate the existence of periodic solutions. The stability of periodic solutions is investigated by using Analogue of Poincar\''{e} Criterion. At last, numerical simulations are given to verify the theoretical analysis.     

具非线性传染率与生物化学控制的害虫管理S-I模型

讨论了具有非线性传染率与脉冲控制的害虫管理S-I传染病模型,此模型考虑的是脉冲投放病虫和喷洒农药.不但得到了系统的所有解的一致完全有界,而且得到了害虫灭绝的边界周期解的全局渐进稳定和系统的一致持久的条件.为实际的害虫管理提供了可靠的理论依据.