Hopf bifurcations in a predator–prey system with a discrete delay and a distributed delay

A delayed Lotka–Volterra two-species predator–prey system with discrete hunting delay and distributed maturation delay for the predator population described by an integral with a strong delay kernel is considered. By linearizing the system at the positive equilibrium and analyzing the associated characteristic equation, the asymptotic stability of the positive equilibrium is investigated and Hopf bifurcations are demonstrated. It is found that under suitable conditions on the parameters the positive equilibrium is asymptotically stable when the hunting delay is less than a certain critical value and unstable when the hunting delay is greater than this critical value. Meanwhile, according to the Hopf bifurcation theorem for functional differential equations (FDEs), we find that the system can also undergo a Hopf bifurcation of nonconstant periodic solution at the positive equilibrium when the hunting delay crosses through a sequence of critical values. In particular, by applying the normal form theory and the center manifold reduction for FDEs, an explicit algorithm determining the direction of Hopf bifurcations and the stability of bifurcating periodic solutions occurring through Hopf bifurcations is given. Finally, to verify our theoretical predictions, some numerical simulations are also included at the end of this paper.     

一类带无选择性捕获和时滞的捕食食饵系统的Hopf分支分析

本文主要研究了一个改进的带时滞和无选择捕获函数的捕食-食饵生态经济系统的稳定性和Hopf分支.利用微分代数系统的稳定性理论和分支理论,得到了系统正平衡点稳定性的条件,以及当时滞τ作为分支参数时系统产生Hopf分支的条件.对Leslie-Gower捕食-食饵模型进行了一定程度的完善,使得建立的模型更符合实际情况,因此得到的结论也更加科学.     

带有非线性收获和妊娠时滞的微分代数捕食者-食饵系统的稳定性及Hopf分支

本文研究了一类同时带有非线性食饵收获和捕食者妊娠时滞的微分代数捕食者-食饵系统的稳定性及Hopf分支问题.利用了分支理论和稳定性理论,以捕食者妊娠时滞作为系统的分支参数,获得了所提出的新系统在正平衡点处系统稳定性的相关判据条件和Hopf分支的产生条件.推广了一般带有线性收获和时滞的微分代数捕食者-食饵系统的结论.     

Dynamics in a diffusive predator–prey system with a constant prey refuge and delay

In this paper, a diffusive predator–prey system with a constant prey refuge and time delay subject to Neumann boundary condition is considered. Local stability and Turing instability of the positive equilibrium are studied. The effect of time delay on the model is also obtained, including locally asymptotical stability and existence of Hopf bifurcation at the positive equilibrium. And the properties of Hopf bifurcation are determined by center manifold theorem and normal form theorem of partial functional differential equations. Some numerical simulations are carried out.     

Stability analysis in a diffusional immunosuppressive infection model with delayed antiviral immune response

In this paper, the diffusion is introduced to an immunosuppressive infection model with delayed antiviral immune response. The direction and stability of Hopf bifurcation are effected by time delay, in the absence of which the positive equilibrium is locally asymptotically stable by means of analyzing eigenvalue spectrum; however, when the time delay increases beyond a threshold, the positive equilibrium loses its stability via the Hopf bifurcation. The stability and direction of the Hopf bifurcation is investigated with the norm form and the center manifold theory. The stability of the Hopf bifurcation leads to the emergence of spatial spiral patterns. Numerical calculations are performed to illustrate our theoretical results. Copyright © 2017 John Wiley & Sons, Ltd.     

Bifurcated periodic solutions in an amensalism system with strong generic delay kernel

The dynamics of an amensalism system with strong generic delay kernel is considered. From a careful analysis of characteristic equation, the local asymptotic stability of the positive equilibrium and the existence of Hopf bifurcation from the positive equilibrium are investigated, where the time delay is used as the bifurcation parameter. Moreover, we apply the normal form theory and the center manifold theorem to study the direction of these Hopf bifurcations and the stability of bifurcated periodic orbits. Finally, numerical simulations supporting the theoretical analysis are also included. Copyright © 2012 John Wiley & Sons, Ltd.     

时滞偏害系统的稳定性和分歧分析

本文讨论了具离散和分布时滞的偏害系统.以时滞作为分歧参数,通过分析原系统在正平衡点处线性化系统的特征方程,获得了正平衡点渐近稳定以及在它周围分歧出周期解的条件.另外,通过使用规范形和中心流形定理,我们获得了Hopf分歧的方向和分歧周期解稳定性的显式算法.最后,数值模拟支持了我们的理论分析.     

Effect of herd shape in a diffusive predator-prey model with time delay

In this paper, we deal with the effect of the shape of herd behavior on the interaction between predator and prey. The model analysis was studied in three parts. The first, The analysis of the system in the absence of spatial diffusion and the time delay, where the local stability of the equilibrium states, the existence of Hopf bifurcation have been investigated. For the second part, the spatiotemporal dynamics introduce by self diffusion was determined, where the existence of Hopf bifurcation, Turing driven instability, Turing-Hopf bifurcation point have been proved. Further, the order of Hopf bifurcation points and regions of the stability of the non trivial equilibrium state was given. In the last part of the paper, we studied the delay effect on the stability of the non trivial equilibrium, where we proved that the delay can lead to the instability of interior equilibrium state, and also the existence of Hopf bifurcation. A numerical simulation was carried out to insure the theoretical results.     

具有捕食者相互残杀项时滞系统的Hopf分支

研究了具有捕食者相互残杀项的时滞系统的Hopf分支,通过选择时滞作为一个分支参数,研究了正平衡点的稳定性和正周期解的Hopf分支.而且通过应用规范型和中心流形的理论,得出了确定分支方向的明确的算法.     

Stability and bifurcation analysis of a nutrient-phytoplankton model with time delay

We proposed a nutrient-phytoplankton interaction model with a discrete and distributed time delay to provide a better understanding of phytoplankton growth dynamics and nutrient-phytoplankton oscillations induced by delay. Standard linear analysis indicated that delay can induce instability of a positive equilibrium via Hopf bifurcation. We derived the conditions guaranteeing the existence of Hopf bifurcation and tracked its direction and the stability of the bifurcating periodic solutions. We also obtained the sufficient conditions for the global asymptotic stability of the unique positive steady state. Numerical analysis in the fully nonlinear regime showed that the stability of the positive equilibrium is sensitive to changes in delay values under select conditions. Numerical results were consistent with results predicted by linear analysis.     

一类具有时滞的捕食——食饵系统的稳定性与Hopf分支

研究了一类具有时滞的捕食—食饵系统,通过分析正平衡点处的特征方程,讨论了系统正平衡点的稳定性;以时滞作为分支参数,应用Hopf分支理论,得到了系统存在Hopf分支的充分条件.     

Stability and Hopf bifurcation of a modified delay predator-prey model with stage structure

In this paper, a modified delay predator-prey model with stage structure is established, which involves the economic factor and internal competition of all the prey and predator populations. By the methods of normal form and characteristic equation, we obtain the stability of the positive equilibrium point and the sufficient condition of the existence of Hopf bifurcation. We analyze the influence of the time delay on the equation and show the occurrence of Hopf bifurcation periodic solution. The simulation gives a visual understanding for the existence and direction of Hopf bifurcation of the model.     

疾病在食饵中传播的具有时滞的捕食被捕食模型的分析

研究了一个疾病在食饵中传播的捕食与被捕食模型.在未引入时滞时,利用Routh-Hurwitz定理证明了正平衡点的局部渐近稳定性.在引入时滞后,主要讨论了正平衡点的稳定性,得到了当经过一系列临界条件时发生Hopf分支.     

Dynamics of a delayed predator–prey model with predator migration

Based on the availability of prey and a simple predator–prey model, we propose a delayed predator–prey model with predator migration to describe biological control. We first study the existence and stability of equilibria. It turns out that backward bifurcation occurs with the migration rate as bifurcation parameter. The stability of the trivial equilibrium and the boundary equilibrium is delay-independent. However, the stability of the positive equilibrium may be delay-dependent. Moreover, delay can switch the stability of the positive equilibrium. When the positive equilibrium loses stability, Hopf bifurcation can occur. The direction and stability of Hopf bifurcation is derived by applying the center manifold method and the normal form theory. The main theoretical results are illustrated with numerical simulations.     

Stability and Hopf bifurcation in a delayed competition system

In this paper, Hopf bifurcation for two-species Lotka–Volterra competition systems with delay dependence is investigated. By choosing the delay as a bifurcation parameter, we prove that the system is stable over a range of the delay and beyond that it is unstable in the limit cycle form, i.e., there are periodic solutions bifurcating out from the positive equilibrium. Our results show that a stable competition system can be destabilized by the introduction of a maturation delay parameter. Further, an explicit algorithm for determining the direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions is derived by using the theory of normal forms and center manifolds, and numerical simulations supporting the theoretical analysis are also given.     

Dynamical Property Analysis of a Delayed Diffusive Predator-prey Model with Fear Effect

In this paper, we study a delayed diffusive predator-prey model with fear effect and Holling II functional response. The stability of the positive equilibrium is investigated. We find that time delay can destabilize the stable equilibrium and induce Hopf bifurcation. Diffusion may lead to Turing instability and inhomogeneous periodic solutions. Through the theory of center manifold and normal form, some detailed formulas for determining the of Hopf bifurcation are presented. Some numerical simulations are also provided.     

含两时滞捕食-被捕食系统的稳定性及分歧

本文研究一类含两相异时滞的捕食－被捕食系统的稳定性及分歧。首先，我们讨论两相异时滞对系统唯一正平衡点的稳定性的影响，通过对系数与时滞有关的特征方程的分析，建立了一种稳定性判别性。其次，将一个时滞看成分歧参数，而另一个看作固定参数，我们证明了该系统具有HOPF分歧特性。最后，我们讨论了分歧解的稳定性。     

Stability Analysis of an Enterprise Competitive Model with Time Delay

A three-dimensional enterprise competitive model with time delay is considered. Where the delay is regarded as bifurcation parameters. By analyzing the corresponding characteristic equation of positive equilibrium,the local stability of positive equilibrium is regarded. By using the normal form method and center manifold theorem, we give the formula for determining the direction of the Hopf bifurcation and the stability of bifurcating periodic solutions. Numerical simulations are shown to illustrate the obtained results.     

Local and global Hopf bifurcation in a neutral population model with age structure

In this paper, an age‐structured population model with the form of neutral functional differential equation is studied. We discuss the stability of the positive equilibrium by analyzing the characteristic equation. Local Hopf bifurcation results are also obtained by choosing the mature delay as bifurcation parameter. On the center manifold, the normal form of the Hopf bifurcation is derived, and explicit formulae for determining the criticality of bifurcation are theoretically given. Moreover, the global continuation of Hopf bifurcating periodic solutions is investigated by using the global Hopf bifurcation theory of neutral equations. Finally, some numerical examples are carried out to support the main results.     

Stability and Hopf bifurcation analysis on a spruce-budworm model with delay

In this paper, the dynamics of a spruce-budworm model with delay is investigated. We show that there exists Hopf bifurcation at the positive equilibrium as the delay increases. Some sufficient conditions for the existence of Hopf bifurcation are obtained by investigating the associated characteristic equation. By using the theory of normal form and center manifold, explicit expression for determining the direction of Hopf bifurcations and the stability of bifurcating periodic solutions are presented.