Dynamics of a delayed predator-prey model with Allee effect and Holling type II functional response

In this paper, a delayed with Holling type II functional response (Beddington-DeAngelis) and Allee effect predator-prey model is considered. The growth of the prey is affected by the parameter $M$, which defines the Allee effect. In addition, the delay $\tau$ also influences the logistic growth of the prey, which can be interpreted as the maturity time or the gestation period. In the study of the characteristic equation, we observe that the delay $\tau$ also depends on the parameter $M$, which affects the dynamics in the prey population. Considering the delay as a bifurcation parameter, the local asymptotic stability of the positive equilibrium is investigated. On the other hand, we find that the system can also suffer a Hopf bifurcation in the positive equilibrium when the delay passes through a sequence of critical values. In particular, we study the direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions, an explicit algorithm is provided applying the normal form theory and center manifold reduction for the functional differential equations. Finally, numerical simulations that support the theoretical analysis are included.     

一类具有时滞和Holling Ⅲ型功能性反应的捕食模型的稳定性和Hopf分支

研究一类具有时滞和Holling Ⅲ型功能性反应的捕食模型的稳定性和Hopf分支.以滞量为参数,得到了系统正平衡点的稳定性和Hopf分支存在的充分条件,给出了确定Hopf分支方向和分支周期解的稳定性的计算公式.     

Ecological complexity and feedback control in a prey–predator system with Holling type III functional response

This article deals with a bioeconomic model of prey–predator system with Holling type III functional response. The dynamical behavior of the system is extensively discussed. Continuous type gestational delay of predators is incorporated in the system to study delay induced instability. It is observed that the system undergoes singularity induced bifurcation at interior equilibrium point when net economic revenue of the system increases through zero. State feedback controller is designed to stabilize the system at positive economic profit. Time delay is considered as a bifurcation parameter to prove the occurrence of Hopf bifurcation phenomenon in the neighborhood of the coexisting equilibrium point. Finally, some numerical simulations are carried out to verify the analytical results and the system is analyzed through graphical illustrations. © 2015 Wiley Periodicals, Inc. Complexity 21: 346–360, 2016     

Positive solutions for a Lotka-Volterra prey-predator model with cross-diffusion and Holling type-Ⅱ functional response

We consider a Lotka-Volterra prey-predator model with cross-diffusion and Holling type-II functional response.The main concern is the existence of positive solutions under the combined effect of cross-diffusion and Holling type-II functional response.Here,a positive solution corresponds to a coexistence state of the model.Firstly,we study the sufficient conditions to ensure the existence of positive solutions by using degree theory and analyze the coexistence region in parameter plane.In addition,we present the uniqueness of positive solutions in one dimension case.Secondly,we study the stability of the trivial and semi-trivial solutions by analyzing the principal eigenvalue of the corresponding linearized system,and then we characterize the stable/unstable regions of semi-trivial solutions in parameter plane.     

Global dynamics of a delayed eco‐epidemiological model with Holling type‐III functional response

In this paper, an eco‐epidemiological model with Holling type‐III functional response and a time delay representing the gestation period of the predators is investigated. In the model, it is assumed that the predator population suffers a transmissible disease. The disease basic reproduction number is obtained. By analyzing the corresponding characteristic equations, the local stability of each of feasible equilibria and the existence of Hopf bifurcations at the disease‐free equilibrium and the endemic‐coexistence equilibrium are established, respectively. By using the persistence theory on infinite dimensional systems, it is proved that if the disease basic reproduction number is greater than unity, the system is permanent. By means of Lyapunov functionals and LaSalle's invariance principle, sufficient conditions are obtained for the global stability of the endemic‐coexistence equilibrium, the disease‐free equilibrium and the predator‐extinction equilibrium of the system, respectively. Copyright © 2013 John Wiley & Sons, Ltd.     

一类具有时滞的捕食模型的稳定性和Hopf分支

研究一类具有时滞和Beddington-DeAngelis功能性反应的捕食模型的稳定性和Hopf分支.以滞量为参数,得到了系统正平衡点的稳定性和Hopf分支存在的充分条件.应用一般泛函微分方程的度理论,研究了该系统的全局Hopf分支的存在性.     

Bifurcation analysis of a generalized Gause model with prey harvesting and a generalized Holling response function of type III

In this paper we study a generalized Gause model with prey harvesting and a generalized Holling response function of type III: . The goal of our study is to give the bifurcation diagram of the model. For this we need to study saddle-node bifurcations, Hopf bifurcation of codimension 1 and 2, heteroclinic bifurcation, and nilpotent saddle bifurcation of codimension 2 and 3. The nilpotent saddle of codimension 3 is the organizing center for the bifurcation diagram. The Hopf bifurcation is studied by means of a generalized Liénard system, and for b=0 we discuss the potential integrability of the system. The nilpotent point of multiplicity 3 occurs with an invariant line and can have a codimension up to 4. But because it occurs with an invariant line, the effective highest codimension is 3. We develop normal forms (in which the invariant line is preserved) for studying of the nilpotent saddle bifurcation. For b=0, the reversibility of the nilpotent saddle is discussed. We study the type of the heteroclinic loop and its cyclicity. The phase portraits of the bifurcations diagram (partially conjectured via the results obtained) allow us to give a biological interpretation of the behavior of the two species.     

Hopf bifurcation analysis in a multiple delayed innovation diffusion model with Holling II functional response

A nonlinear mathematical model with Holling II functional response describing the dynamics of nonadopter and adopters population in a stage structured innovation diffusion model, which incorporates the evaluation stage (multiple delays), is proposed. Firstly, we study the stability and the existence of periodic solutions via Hopf bifurcation with respect to both delays at the positive equilibrium by analyzing the distribution of the roots of the corresponding exponential characteristic equation obtained through the variational matrix. The direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions are determined with the help of normal form theory and center manifold theorem. Meanwhile, various cases are discussed to examine the effect of different delays on the stability of delayed innovation diffusion system and are also established numerically. It is also observed that the cumulative density of external influences has a significant role in developing maturity stage (adoption stage) in the system. Finally, numerical simulations are carried out to support and supplement the analytical findings.     

Bifurcation analysis of a predator–prey model with memory-based diffusion

In this paper, we investigate the predator–prey model equipped with Fickian diffusion and memory-based diffusion of predators. The stability and bifurcation analysis explores the impacts of the memory-based diffusion and the averaged memory period on the dynamics near the positive steady state. Specifically, when the memory-based diffusion coefficient is less than a critical value, we show that the stability of the positive steady state can be destabilized as the average memory period increases, which leads to the occurrence of Hopf bifurcations. Moreover, we also analyze the bifurcation properties using the central manifold theorem and normal form theory. This allows us to prove the existence of stable spatially inhomogeneous periodic solutions arising from Hopf bifurcation. In addition, the sufficient and necessary conditions for the occurrence of stability switches are also provided.     

Dynamics and bifurcations of a modified Leslie‐Gower–type model considering a Beddington‐DeAngelis functional response

In this paper, a planar system of ordinary differential equations is considered, which is a modified Leslie‐Gower model, considering a Beddington‐DeAngelis functional response. It generates a complex dynamics of the predator‐prey interactions according to the associated parameters. From the system obtained, we characterize all the equilibria and its local behavior, and the existence of a trapping set is proved. We describe different types of bifurcations (such as Hopf, Bogdanov‐Takens, and homoclinic bifurcation), and the existence of limit cycles is shown. Analytic proofs are provided for all results. Ecological implications and a set of numerical simulations supporting the mathematical results are also presented.     

Stability and bifurcation analysis of a viral infection model with delayed immune response

In this paper, we study a viral infection model with an immunity time delay accounting for the time between the immune system touching antigenic stimulation and generating CTLs. By calculation, we derive two thresholds to determine the global dynamics of the model, i.e., the reproduction number for viral infection $R_{0}$ and for CTL immune response $R_{1}$. By analyzing the characteristic equation, the local stability of each feasible equilibrium is discussed. Furthermore, the existence of Hopf bifurcation at the CTL-activated infection equilibrium is also studied. By constructing suitable Lyapunov functionals, we prove that when $R_{0}\leq1$, the infection-free equilibrium is globally asymptotically stable; when $R_{0}>1$ and $R_{1}\leq1$, the CTL-inactivated infection equilibrium is globally asymptotically stable; Numerical simulation is carried out to illustrate the main results in the end.     

Stability and Hopf bifurcation in a viral infection model with nonlinear incidence rate and delayed immune response

A viral infection model with nonlinear incidence rate and delayed immune response is investigated. It is shown that if the basic reproduction ratio of the virus is less than unity, the infection-free equilibrium is globally asymptotically stable. By analyzing the characteristic equation, the local stability of the chronic infection equilibrium of the system is discussed. Furthermore, the existence of Hopf bifurcations at the chronic infection equilibrium is also studied. By means of an iteration technique, sufficient conditions are obtained for the global attractiveness of the chronic infection equilibrium. Numerical simulations are carried out to illustrate the main results.     

Bifurcations in a diffusive predator–prey model with predator saturation and competition response

A diffusive predator–prey model with predator saturation and competition response subject to homogeneous Neumann boundary conditions is considered in this paper. We find that the spatially homogeneous and non‐homogeneous periodic solutions through all parameters of the system are spatially homogeneous. To verify our theoretical results, some numerical simulations are also carried out. Copyright © 2014 John Wiley & Sons, Ltd.     

Dynamics of a ratio‐dependent stage‐structured predator‐prey model with delay

In this paper, we investigate the dynamics of a time‐delay ratio‐dependent predator‐prey model with stage structure for the predator. This predator‐prey system conforms to the realistically biological environment. The existence and stability of the positive equilibrium are thoroughly analyzed, and the sufficient and necessary conditions for the stability and instability of the positive equilibrium are obtained for the case without delay. Then, the influence of delay on the dynamics of the system is investigated using the geometric criterion developed by Beretta and Kuang. 26 We show that the positive steady state can be destabilized through a Hopf bifurcation and there exist stability switches under some conditions. The formulas determining the direction and the stability of Hopf bifurcations are explicitly derived by using the center manifold reduction and normal form theory. Finally, some numerical simulations are performed to illustrate and expand our theoretical results.     

Stationary distribution and extinction of a stochastic one-prey two-predator model with Holling type II functional response

In this article, we investigate a stochastic one-prey two-predator model with Holling type II functional response. We first establish sufficient conditions for persistence and extinction of prey and predator populations, then by constructing a suitable stochastic Lyapunov function, we establish sharp sufficient criteria for the existence of a unique ergodic stationary distribution of the positive solutions to the model. The results show that the smaller white noise can ensure the persistence of prey and predator populations while the larger white noise can lead to the extinction of prey and predator populations.     

Global stability and Hopf bifurcation of a delayed eco‐epidemiological model with Holling type II functional response

In this paper, a delayed eco‐epidemiological model with Holling type II functional response is investigated. By analyzing corresponding characteristic equations, the local stability of each of the feasible equilibria and the existence of Hopf bifurcations at the disease‐free equilibrium, the susceptible predator‐free equilibrium and the endemic‐coexistence equilibrium are established, respectively. By means of Lyapunov functionals and LaSalle's invariance principle, sufficient conditions are derived for the global stability of the endemic‐coexistence equilibrium, the disease‐free equilibrium, the susceptible predator‐free equilibrium and the predator‐extinction equilibrium of the system, respectively. Numerical simulations are carried out to illustrate the theoretical results. Copyright © 2014 John Wiley & Sons, Ltd.     

Stability and Hopf bifurcation of a delayed predator-prey system with nonlocal competition and herd behaviour

In this paper, we investigate the stability and Hopf bifurcation of a diffusive predator-prey system with herd behaviour. The model is described by introducing both time delay and nonlocal prey intraspecific competition. Compared to the model without time delay, or without nonlocal competition, thanks to the together action of time delay and nonlocal competition, we prove that the first critical value of Hopf bifurcation may be homogenous or non-homogeneous. We also show that a double-Hopf bifurcation occurs at the intersection point of the homogenous and non-homogeneous Hopf bifurcation curves. Furthermore, by the computation of normal forms for the system near equilibria, we investigate the stability and direction of Hopf bifurcation. Numerical simulations also show that the spatially homogeneous and non-homogeneous periodic patters.     

Stability and Hopf bifurcation of a diffusive predator‐prey model with hyperbolic mortality

The dynamics of a reaction‐diffusion predator‐prey model with hyperbolic mortality and Holling type II response effect is considered. The stability of the positive equilibrium and the existence of Hopf bifurcation are investigated by analyzing the distribution of eigenvalues without diffusion. We also study the spatially homogeneous and nonhomogeneous periodic solutions through all parameters of the system which are spatially homogeneous. To verify our theoretical results, some numerical simulations are also presented. © 2015 Wiley Periodicals, Inc. Complexity 21: 34–43, 2016     

Coexistence states of a Holling type-II predator-prey system

This paper characterize the existence of coexistence states to a reaction-diffusion predator-prey model with Holling type-II functional response subject to Dirichlet boundary conditions. We find the necessary and sufficient conditions for existence of coexistence states by fixed point index theory and bifurcation theory.