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1.
To study the mechanisms that govern the coding of temporal features of complex sound signals, responses of single neurons located in the dorsal nucleus of the medulla oblongata (the cochlear nucleus) of a curarized grass frog (Rana temporaria) to pure tone bursts and amplitude modulated tone bursts with a modulation frequency of 20 Hz and modulation depths of 10 and 80% were recorded. The carrier frequency was equal to the characteristic frequency of a neuron, the average signal level was 20–30 dB above the threshold, and the signal duration was equal to ten full modulation periods. Of the 133 neurons studied, 129 neurons responded to 80% modulated tone bursts by discharges that were phase-locked with the envelope waveform. At this modulation depth, the best phase locking was observed for neurons with the phasic type of response to tone bursts. For tonic neurons with low characteristic frequencies, along with the reproduction of the modulation, phase locking with the carrier frequency of the signal was observed. At 10% amplitude modulation, phasic neurons usually responded to only the onset of a tone burst. Almost all tonic units showed a tendency to reproduce the envelope, although the efficiency of the reproduction was low, and for half of these neurons, it was below the reliability limit. Some neurons exhibited a more efficient reproduction of the weak modulation. For almost half of the neurons, a reliable improvement was observed in the phase locking of the response during the tone burst presentation (from the first to the tenth modulation period). The cooperative histogram of a set of neurons responding to 10% modulated tone bursts within narrow ranges of frequencies and intensities retains the information on the dynamics of the envelope variation. The data are compared with the results obtained from the study of the responses to similar signals in the acoustic midbrain center of the same object and also with the psychophysical effect of a differential sensitivity increase in the process of adaptation.  相似文献   

2.
A model is presented which calculates the intrinsic envelope power of a bandpass noise carrier within the passband of a hypothetical modulation filter tuned to a specific modulation frequency. Model predictions are compared to experimentally obtained amplitude modulation (AM) detection thresholds. In experiment 1, thresholds for modulation rates of 5, 25, and 100 Hz imposed on a bandpass Gaussian noise carrier with a fixed upper cutoff frequency of 6 kHz and a bandwidth in the range from 1 to 6000 Hz were obtained. In experiment 2, three noises with different spectra of the intrinsic fluctuations served as the carrier: Gaussian noise, multiplied noise, and low-noise noise. In each case, the carrier was spectrally centered at 5 kHz and had a bandwidth of 50 Hz. The AM detection thresholds were obtained for modulation frequencies of 10, 20, 30, 50, 70, and 100 Hz. The intrinsic envelope power of the carrier at the output of the modulation filter tuned to the signal modulation frequency appears to provide a good estimate for AM detection threshold. The results are compared with predictions on the basis of the more complex auditory processing model by Dau et al.  相似文献   

3.
A binaural unmasking of a tone component that is present in an amplitude-time noise envelope of a high-frequency signal is studied. The signal has the form of a sinusoidal carrier of frequency 2000–5000 Hz amplitude modulated by a low-frequency signal. The modulating function is a mixture of a 300-Hz tone (interaurally inphase or antiphase) and a dichotic masking noise within 0–400 Hz, this mixture being subjected to a half-wave linear rectification. The listener has to detect the rhythmic component in the modulating noise function. It is shown that, under the aforementioned conditions, the binaural difference in masking levels grows up to 25 dB with increasing carrier frequency but drastically decreases in the case of a masking of the low-frequency part of the basilar membrane in the vicinity of 300 Hz. The lateralization based on the interaural phase of a 100% amplitude modulation by a 300-Hz tone at a carrier frequency within 2000 to 5000 Hz also drastically decreases (in our experiments) when the low-frequency part of the basilar membrane is masked.  相似文献   

4.
孟庆林  原猛  牟宏宇  陈友元  冯海泓 《物理学报》2012,61(16):164302-164302
通过心理物理实验探讨了包络调制率(<300 Hz)和纯音载波频率(<8 kHz)对听觉时间调制检测能力的影响. 测试信号为以纯音为载波的正弦幅度调制信号, 采用二选一强迫选择法和自适应调整步长的心理物理实验方法, 测试得到不同载波频率条件下的时间调制传递函数. 实验结果表明, 包络调制率和载波频率均会对听觉的时间调制检测能力产生影响. 当载波频率低于2 kHz时, 人耳的检测能力与调制率呈单调递增趋势;当载波频率高于3.5 kHz时, 检测能力也会受到调制率的显著影响, 但没有显著的单调变化趋势. 当调制率在10-100 Hz之间时, 检测能力不随载波频率明显变化;当调制率在150-300 Hz之间时, 调制检测能力随着载波频率上升而下降, 在载波频率达到3.5 kHz时, 调制检测能力不随载波频率显著改变.  相似文献   

5.
Steady-state evoked potential responses were measured to binaural amplitude-modulated (AM) and combined amplitude- and frequency-modulated (AM/FM) tones. For awake subjects, AM/FM tones produced larger amplitude responses than did AM tones. Awake and sleeping responses to 30-dB HL AM/FM tones were compared. Response amplitudes were lower during sleep and the extent to which they differed from awake amplitudes was dependent on both carrier and modulation frequencies. Background EEG noise at the stimulus modulation frequency was also reduced during sleep and varied with modulation frequency. A detection efficiency function was used to indicate the modulation frequencies likely to be most suitable for electrical estimation of behavioral threshold. In awake subjects, for all carrier frequencies tested, detection efficiency was highest at a modulation frequency of 45 Hz. In sleeping subjects, the modulation frequency regions of highest efficiency varied with carrier frequency. For carrier frequencies of 250 Hz, 500 Hz, and 1 kHz, the highest efficiencies were found in two modulation frequency regions centered on 45 and 90 Hz. For 2 and 4 kHz, the highest efficiencies were at modulation frequencies above 70 Hz. Sleep stage affected both response amplitude and background EEG noise in a manner that depended on modulation frequency. The results of this study suggest that, for sleeping subjects, modulation frequencies above 70 Hz may be best when using steady-state potentials for hearing threshold estimation.  相似文献   

6.
Spectro-temporal processing in the envelope-frequency domain   总被引:1,自引:0,他引:1  
The frequency selectivity for amplitude modulation applied to tonal carriers and the role of beats between modulators in modulation masking were studied. Beats between the masker and signal modulation as well as intrinsic envelope fluctuations of narrow-band-noise modulators are characterized by fluctuations in the "second-order" envelope (referred to as the "venelope" in the following). In experiment 1, masked threshold patterns (MTPs), representing signal modulation threshold as a function of masker-modulation frequency, were obtained for signal-modulation frequencies of 4, 16, and 64 Hz in the presence of a narrow-band-noise masker modulation, both applied to the same sinusoidal carrier. Carrier frequencies of 1.4, 2.8, and 5.5 kHz were used. The shape and relative bandwidth of the MTPs were found to be independent of the signal-modulation frequency and the carrier frequency. Experiment 2 investigated the extent to which the detection of beats between signal and masker modulation is involved in tone-in-noise (TN), noise-in-tone (NT), and tone-in-tone (TT) modulation masking, whereby the TN condition was similar to the one used in the first experiment. A signal-modulation frequency of 64 Hz, applied to a 2.8-kHz carrier, was tested. Thresholds in the NT condition were always lower than in the TN condition, analogous to the masking effects known from corresponding experiments in the audio-frequency domain. TT masking conditions generally produced the lowest thresholds and were strongly influenced by the detection of beats between the signal and the masker modulation. In experiment 3, TT masked-threshold patterns were obtained in the presence of an additional sinusoidal masker at the beat frequency. Signal-modulation frequencies of 32, 64, and 128 Hz, applied to a 2.8-kHz carrier, were used. It was found that the presence of an additional modulation at the beat frequency hampered the subject's ability to detect the envelope beats and raised thresholds up to a level comparable to that found in the TN condition. The results of the current study suggest that (i) venelope fluctuations play a similar role in modulation masking as envelope fluctuations do in spectral masking, and (ii) envelope and venelope fluctuations are processed by a common mechanism. To interpret the empirical findings, a general model structure for the processing of envelope and venelope fluctuations is proposed.  相似文献   

7.
Temporal modulation transfer functions (TMTFs) were measured for detection of monaural sinusoidal amplitude modulation and dynamically varying interaural level differences for a single set of listeners. For the interaural TMTFs, thresholds are the modulation depths at which listeners can just discriminate interaural envelope-phase differences of 0 and 180 degrees. A 5-kHz pure tone and narrowband noises, 30- and 300-Hz wide centered at 5 kHz, were used as carriers. In the interaural conditions, the noise carriers were either diotic or interaurally uncorrelated. The interaural TMTFs with tonal and diotic noise carriers exhibited a low-pass characteristic but the cutoff frequencies changed nonmonotonically with increasing bandwidth. The interaural TMTFs for the tonal carrier began rolling off approximately a half-octave lower than the tonal monaural TMTF (approximately 80 Hz vs approximately 120 Hz). Monaural TMTFs obtained with noise carriers showed effects attributable to masking of the signal modulation by intrinsic fluctuations of the carrier. In the interaural task with dichotic noise carriers, similar masking due to the interaural carrier fluctuations was observed. Although the mechanisms responsible for differences between the monaural and interaural TMTFs are unknown, the lower binaural TMTF cutoff frequency suggests that binaural processing exhibits greater temporal limitation than monaural processing.  相似文献   

8.
These experiments were designed to examine the mechanism of detection of phase disparity in the envelopes of two sinusoidally amplitude-modulated (AM) sinusoids. Specifically, they were performed to determine whether detection of envelope phase disparity was consistent with processing within a single channel in which the AM tones were simply added. In the first condition, with an 8-Hz modulation frequency, phase-disparity thresholds increased sharply with an initial increase in separation of the carrier frequencies. They then remained approximately constant when the separation was an octave or above. In the second condition, with carrier pairs of 1 and 2 kHz or 1 and 3.2 kHz and a modulation frequency of 8 Hz, thresholds were little affected as the level of one carrier was decreased relative to the other. With a modulation frequency of 128 Hz, for most subjects there was more of an effect of level disparity on thresholds. In the third condition, when the modulation frequency was 8 Hz, subjects showed relatively constant thresholds whether the signals were presented monotically, dichotically, or dichotically with low- and high-pass noise. Dichotic thresholds were typically higher than monotic when the modulation frequency was 128 Hz. These results suggest that it is not necessary to have information available within a single additive channel to detect envelope phase disparity. In certain circumstances, a comparison across channels may be used to detect such disparities.  相似文献   

9.
The present study investigates the nature of spectral envelope perception using a spectral modulation detection task in which sinusoidal spectral modulation is superimposed upon a noise carrier. The principal goal of this study is to characterize spectral envelope perception in terms of the influence of modulation frequency (cycles/octave), carrier bandwidth (octaves), and carrier frequency region (defined by lower and upper cutoff frequencies in Hz). Spectral modulation detection thresholds measured as a function of spectral modulation frequency result in a spectral modulation transfer function (SMTF). The general form of the SMTF is bandpass in nature, with a minimum modulation detection threshold in the region between 2 to 4 cycles/octave. SMTFs are not strongly dependent on carrier bandwidth (ranging from 1 to 6 octaves) or carrier frequency region (ranging from 200 to 12 800 Hz), with the exception of carrier bands restricted to very low audio frequencies (e.g., 200-400 Hz). Spectral modulation detection thresholds do not depend on the presence of random level variations or random modulation phase across intervals. The SMTFs reported here and associated excitation pattern computations are considered in terms of a linear systems approach to spectral envelope perception and potential underlying mechanisms for the perception of spectral features.  相似文献   

10.
Experiments were performed to determine under what conditions quasi-frequency-modulated (QFM) noise and random-sideband noise are suitable comparisons for AM noise in measuring a temporal modulation transfer function (TMTF). Thresholds were measured for discrimination of QFM from random-sideband noise and AM from QFM noise as a function of sideband separation. In the first experiment, the upper spectral edge of the noise stimuli was at 2400 Hz and the bandwidth was 1600 Hz. For sideband separations up to 256 Hz, at threshold sideband levels for discriminating AM from QFM noise, QFM was indiscriminable from random-sideband noise. For the largest sideband separation used (512 Hz), listeners may have used within-stimulus envelope correlation in the QFM noise to discriminate it from the random-sideband noise. Results when stimulus bandwidth was varied suggest that listeners were able to use this cue when the carrier was wider than a critical band, and the sideband separation approached the carrier bandwidth. Within-stimulus envelope correlation was also present in AM noise, and thus QFM noise was a suitable comparison because it made this cue unusable and forced listeners to use across-stimulus envelope differences. When the carrier bandwidth was less than a critical band or was wideband, QFM noise and random-sideband noise were equally suitable comparisons for AM noise. When discrimination thresholds for QFM and random-sideband noise were converted to modulation depth and modulation frequency, they were nearly identical to those for discrimination of AM from QFM noise, suggesting that listeners were using amplitude modulation cues in both cases.  相似文献   

11.
It has been proposed that the detection of frequency modulation (FM) of sinusoidal carriers can be mediated by two mechanisms; a place mechanism based on FM-induced amplitude modulation (AM) in the excitation pattern, and a temporal mechanism based on phase locking in the auditory nerve. The temporal mechanism appears to be "sluggish" and does not play a role for FM rates above about 10 Hz. It also does not play a role for high carrier frequencies (above about 5 kHz). This experiment provided a further test of the hypothesis that the effectiveness of the temporal mechanism depends upon the time spent close to frequency extremes during the modulation cycle. Psychometric functions for the detection of AM and FM were measured for two carrier frequencies, 1 and 6 kHz. The modulation waveform was quasitrapezoidal. Within each modulation period, P, a time Tss was spent at each extreme of frequency or amplitude. The transitions between the extremes, with duration Ttrans had the form of a half-cycle of a cosine function. The modulation rate was 2, 5, 10, or 20 Hz, giving values of P of 500, 200, 100, and 50 ms. TSS varied from 0 ms (sinusoidal modulation) up to 160, 80, 40, or 20 ms, for rates of 2, 5, 10, and 20 Hz, respectively. The detectability of AM was not greatly affected by modulation rate or by the value of TSS, except for a slight improvement with increasing TSS for the lowest modulation rates; this was true for both carrier frequencies. For FM of the 6-kHz carrier, the pattern of results was similar to that found for AM, which is consistent with an excitation-pattern model of FM detection. For FM of the 1-kHz carrier, performance improved markedly with increasing TSS, especially for the lower FM rates; there was no change in performance with TSS for the 20-Hz modulation rate. The results are consistent with the idea that detection of FM of a 1-kHz carrier is partly mediated by a sluggish temporal mechanism. That mechanism benefits from greater time spent at frequency extremes of the modulation cycle for rates up to 10 Hz.  相似文献   

12.
The effect of temporal asymmetry on amplitude modulation detection was studied using sawtooth modulators with rising (ramped) or falling (damped) temporal envelopes within each period of modulation. For pure-tone carriers, damped modulation was more detectable than ramped modulation for a 5-kHz carrier (by a threshold difference of 3.2 dB on average) but not for a 1-kHz carrier. The threshold difference obtained at 5 kHz between the ramped and damped modulators was consistent across modulation rates (8-128 Hz). This carrier frequency dependence suggests that the effect of temporally asymmetry on modulation detection originates from envelope-based, within-channel mechanisms.  相似文献   

13.
Detectability of a tonal signal added to a tonal masker increases with increasing duration ("temporal integration"), up to some maximum duration. Initially assumed to be some form of energy integration over time, this phenomenon is now often described as the result of a statistical "multiple looks" process. For continuous maskers, listeners may also use a mechanism sensitive to changes in stimulus intensity, possibly a result of inherent sensitivity to amplitude modulation (AM). In order to examine this hypothesis, change detection was investigated in the presence of AM maskers presented at either the same carrier frequency as the target signal or at a distant frequency. The results are compatible with the hypothesis that listeners detect intensity increments by using change-detection mechanisms (modeled here as the outputs of a bank of modulation filters) sensitive to envelope modulation at both low (4-16 Hz) and high (around 100 Hz) rates. AM masking occurred even when the masker was at a carrier frequency more than two octaves above that of the signal to be detected. This finding is also compatible with the hypothesis that similar mechanisms underlie sensitivity to AM (where across-frequency masking is commonly shown) and detection of intensity increments.  相似文献   

14.
Detection thresholds were measured for a sinusoidal modulation applied to the modulation depth of a sinusoidally amplitude-modulated (SAM) white noise carrier as a function of the frequency of the modulation applied to the modulation depth (referred to as f'm). The SAM noise acted therefore as a "carrier" stimulus of frequency fm, and sinusoidal modulation of the SAM-noise modulation depth generated two additional components in the modulation spectrum: fm-f'm and fm+f'm. The tracking variable was the modulation depth of the sinusoidal variation applied to the "carrier" modulation depth. The resulting "second-order" temporal modulation transfer functions (TMTFs) measured on four listeners for "carrier" modulation frequencies fm of 16, 64, and 256 Hz display a low-pass segment followed by a plateau. This indicates that sensitivity to fluctuations in the strength of amplitude modulation is best for fluctuation rates f'm below about 2-4 Hz when using broadband noise carriers. Measurements of masked modulation detection thresholds for the lower and upper modulation sideband suggest that this capacity is possibly related to the detection of a beat in the sound's temporal envelope. The results appear qualitatively consistent with the predictions of an envelope detector model consisting of a low-pass filtering stage followed by a decision stage. Unlike listeners' performance, a modulation filterbank model using Q values > or = 2 should predict that second-order modulation detection thresholds should decrease at high values of f'm due to the spectral resolution of the modulation sidebands (in the modulation domain). This suggests that, if such modulation filters do exist, their selectivity is poor. In the latter case, the Q value of modulation filters would have to be less than 2. This estimate of modulation filter selectivity is consistent with the results of a previous study using a modulation-masking paradigm [S. D. Ewert and T. Dau, J. Acoust. Soc. Am. 108, 1181-1196 (2000)].  相似文献   

15.
To better understand the processing of complex high-frequency sounds, modulation-detection thresholds were measured for sinusoidal frequency modulation (SFM), quasi-frequency modulation (QFM), sinusoidal amplitude modulation (SAM), and random-phase FM (RPFM). At the lowest modulation frequency (5 Hz) modulation thresholds expressed as AM depth were similar for RPFM, SAM and QFM suggesting the predominance of envelope cues. At the higher modulation frequencies (20 and 40 Hz) thresholds expressed as total frequency excursions were similar for SFM and QFM suggesting a common mechanism, one perhaps based on single-channel FM-to-AM conversion or on a multi-channel place mechanism. The fact that the nominal envelopes of SFM and QFM are different (SFM has a flat envelope), seems to preclude processing based on the envelope of the external stimulus. Also, given the 4-kHz carrier and the similarity to previously published results obtained with a 1-kHz carrier, processing based on temporally-coded fine structure for all four types of modulation appears unlikely.  相似文献   

16.
Gupta KK  Onodera N 《Optics letters》2005,30(17):2221-2223
We demonstrate a new method of optical pulse generation in regeneratively mode-locked fiber ring lasers (RML-FRLs). The method is based on generating dominant longitudinal modes in the fiber ring cavity by means of a composite cavity structure and their intermode beating at the photodetector incorporated in the regenerative feedback loop. The beat signal is then used as a modulation signal to generate optical pulses in a regenerative mode-locking scheme, thereby eliminating the requirement for a high-Q rf bandpass filter in a conventional RML-FRL. Optical pulses with a repetition frequency of 3.6 GHz have been generated successfully with a supermode noise suppression of more than 48 dB and a low phase noise of -85 dBc/Hz at 10 kHz offset from the carrier frequency.  相似文献   

17.
This paper is concerned with modulation and beat detection for sinusoidal carriers. In the first experiment, temporal modulation transfer functions (TMTFs) were measured for carrier frequencies between 1 and 10 kHz. Modulation rates covered the range from 10 Hz to about the rate equaling the critical bandwidth at the carrier frequency. In experiment 2, TMTFs for three carrier frequencies were obtained as a function of the carrier level. In the final experiment, thresholds for the detection of either the lower or the upper modulation sideband (beat detection) were measured for "carrier" frequencies of 5 and 10 kHz, using the same range of modulation rates as in experiment 1. The TMTFs for carrier frequencies of 2 kHz and higher remained flat up to a modulation rate of about 100-130 Hz and had similar values across carrier frequencies. For higher rates, modulation thresholds initially increased and then decreased rapidly, reflecting the subjects' ability to resolve the sidebands spectrally. Detection thresholds generally improved with increasing carrier level, but large variations in the exact level dependence were observed, across subjects as well as across carrier frequencies. For beat rates up to about 70 Hz (at 5 kHz) and 100 Hz (at 10 kHz), beat detection thresholds were the same for the upper and the lower sidebands and were about 6 dB higher than the level per sideband at the modulation-detection threshold. At higher rates the threshold for both sidebands increased, but the increase was larger for the lower sideband. This reflects an asymmetry in masking with more masking towards lower frequencies. Only at rates well beyond the maximum of the TMTF did detection for the lower sideband start to be better than that for the upper sideband. The asymmetry at intermediate frequency separations can be explained by assuming that detection always takes place in filters centered above the stimulus spectrum. The shape of the TMTF and the beat-detection data reflects a limitation in resolving fast amplitude variations, which must occur central to the inner-ear filtering. Its characteristic resembles that of a first-order low-pass filter with a cutoff frequency of about 150 Hz.  相似文献   

18.
Estimates of auditory temporal resolution were obtained from normal chinchillas using sinusoidally amplitude modulated noise. Afterwards, the animals were exposed to noise whose bandwidth was progressively increased toward the low frequencies in octave steps. The first exposure was to an octave band of noise centered at 8 kHz. Three additional octave bands of noise were subsequently added to the original exposure in order to progressively increase the extent of the high-frequency hearing loss. The first exposure produced a temporary hearing loss of 50 to 60 dB near 8 kHz and elevated the amplitude modulation thresholds primarily at intermediate (128 Hz) modulation frequencies. Successive noise exposures extended the temporary hearing loss toward lower frequencies, but there was little further deterioration in the amplitude modulation function until the last exposure when the hearing loss spread to 1 kHz. The degradation in the amplitude modulation function observed after the last exposure, however, was due to a reduction in the sensation level of the test signal rather than to a decrease in the hearing bandwidth. The results of this study suggest that the high-frequency regions of the cochlea may be important for temporal resolution.  相似文献   

19.
The present study sought to establish whether speech recognition can be disrupted by the presence of amplitude modulation (AM) at a remote spectral region, and whether that disruption depends upon the rate of AM. The goal was to determine whether this paradigm could be used to examine which modulation frequencies in the speech envelope are most important for speech recognition. Consonant identification for a band of speech located in either the low- or high-frequency region was measured in the presence of a band of noise located in the opposite frequency region. The noise was either unmodulated or amplitude modulated by a sinusoid, a band of noise with a fixed absolute bandwidth, or a band of noise with a fixed relative bandwidth. The frequency of the modulator was 4, 16, 32, or 64 Hz. Small amounts of modulation interference were observed for all modulator types, irrespective of the location of the speech band. More important, the interference depended on modulation frequency, clearly supporting the existence of selectivity of modulation interference with speech stimuli. Overall, the results suggest a primary role of envelope fluctuations around 4 and 16 Hz without excluding the possibility of a contribution by faster rates.  相似文献   

20.
The addition of a signal in the N0Sπ binaural configuration gives rise to fluctuations in interaural phase and amplitude. Sensitivity to these individual cues was measured by applying sinusoidal amplitude modulation (AM) or quasi-frequency modulation (QFM) to a band of noise. Discrimination between interaurally in-phase and out-of-phase modulation was measured using an adaptive task for narrow bands of noise at center frequencies from 250 to 1500 Hz, for modulation rates of 2-40 Hz, and with or without flanking bands of diotic noise. Discrimination thresholds increased steeply for QFM with increasing center frequency, but increased only modestly for AM, and mainly for modulation rates below 10 Hz. Flanking bands of noise increased thresholds for AM, but had no consistent effect for QFM. The results suggest that two underlying mechanisms may support binaural unmasking: one most sensitive to interaural amplitude modulations that is susceptible to across-frequency interference, and a second, most sensitive to interaural phase modulations that is immune to such effects.  相似文献   

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