Influence of predator mutual interference and prey refuge on Lotka–Volterra predator–prey dynamics

A Lotka–Volterra predator–prey model incorporating a constant number of prey using refuges and mutual interference for predator species is presented. By applying the divergency criterion and theories on exceptional directions and normal sectors, we show that the interior equilibrium is always globally asymptotically stable and two boundary equilibria are both saddle points. Our results indicate that prey refuge has no influence on the coexistence of predator and prey species of the considered model under the effects of mutual interference for predator species, which differently from the conclusion without predator mutual interference, thus improving some known ones. Numerical simulations are performed to illustrate the validity of our results.     

PERMANENCE OF A NONLINEAR PERIODIC PREDATOR-PREY SYSTEM WITH PREY DISPERSAL AND PREDATOR DENSITY-INDEPENDENCE

In this paper,a set of suffcient conditions which ensure the permanence of a nonlinear periodic predator-prey system with prey dispersal and predator density-independence are obtained,where the prey species can disperse among n patches,while the density-independent predator is confined to one of the patches and cannot disperse. Our results generalize some known results.     

On a nonlinear nonautonomous predator–prey model with diffusion and distributed delay

In this paper, a nonlinear nonautonomous predator–prey model with diffusion and continuous distributed delay is studied, where all the parameters are time-dependent. The system, which is composed of two patches, has two species: the prey can diffuse between two patches, but the predator is confined to one patch. We first discuss the uniform persistence and global asymptotic stability of the model; after that, by constructing a suitable Lyapunov functional, some sufficient conditions for the existence of a unique almost periodic solution of the system are obtained. An example shows the feasibility of our main results.     

A safe harbor can protect an endangered species from its predators

The objective is the study of the dynamics of a prey–predator model where the prey species can migrate between two patches. The specialist predator is confined to the first patch, where it consumes the prey following the simple law of mass action. The prey is further “endangered” in that it suffers from the strong Allee effect, assumed to occur due to the lowering of successful matings. In the second patch the prey grows logistically. The model is formulated in a comprehensive way so as to include specialist as well as generalist predators, as a continuum of possible behaviors. This model described by a set of three ordinary differential equation is an extension of some previous models proposed and analysed in the literature on metapopulation models. The following analysis issues will be addressed: boundedness of the solution, equilibrium feasibility and stability, and dynamic behaviour dependency of the population and environmental parameters. Three types for both equilibria and limit cycles are possible: trivial, predator-free and coexistence. Classical analysis techniques are used and also theoretical and numerical bifurcation analysis. Besides the well-known local bifurcations, also a homoclinic connection as a global bifurcation is calculated. In view of the difficulty in the analysis, only the specialist case will be analysed. The obtained results indicate that the safe harbor can protect the endangered species under certain parametric restrictions.

     

Nonconstant Positive Steady States and Pattern Formation of 1D Prey-Taxis Systems

Prey-taxis is the process that predators move preferentially toward patches with highest density of prey. It is well known to have an important role in biological control and the maintenance of biodiversity. To model the coexistence and spatial distributions of predator and prey species, this paper concerns nonconstant positive steady states of a wide class of prey-taxis systems with general functional responses over 1D domain. Linearized stability of the positive equilibrium is analyzed to show that prey-taxis destabilizes prey–predator homogeneity when prey repulsion (e.g., due to volume-filling effect in predator species or group defense in prey species) is present, and prey-taxis stabilizes the homogeneity otherwise. Then, we investigate the existence and stability of nonconstant positive steady states to the system through rigorous bifurcation analysis. Moreover, we provide detailed and thorough calculations to determine properties such as pitchfork and turning direction of the local branches. Our stability results also provide a stable wave mode selection mechanism for thee reaction–advection–diffusion systems including prey-taxis models considered in this paper. Finally, we provide numerical studies of prey-taxis systems with Holling–Tanner kinetics to illustrate and support our theoretical findings. Our numerical simulations demonstrate that the \(2\times 2\) prey-taxis system is able to model the formation and evolution of various striking patterns, such as spikes, periodic oscillations, and coarsening even when the domain is one-dimensional. These dynamics can model the coexistence and spatial distributions of interacting prey and predator species. We also give some insights on how system parameters influence pattern formation in these models.     

An impulsive periodic predator–prey system with Holling type III functional response and diffusion

This paper studies an impulsive two species periodic predator–prey Lotka–Volterra type dispersal system with Holling type III functional response in a patchy environment, in which the prey species can disperse among n different patches, but the predator species is confined to one patch and cannot disperse. Conditions for the permanence and extinction of the predator–prey system, and for the existence of a unique globally stable periodic solution are established. Numerical examples are shown to verify the validity of our results.     

Existence and global stability of positive periodic solutions for predator–prey system with infinite delay and diffusion

In this paper, we study a periodic predator–prey system with Holling type III functional response, in which the prey species can diffuse among two patches but the predator is confined in one patch. By using the continuation theorem of coincidence degree theory and Lyapunov functional, some sufficient conditions are obtained.     

Dynamics of predator–prey models with a strong Allee effect on the prey and predator-dependent trophic functions

The complex dynamics of a two-trophic chain are investigated. The chain is described by a general predator–prey system, in which the prey growth rate and the trophic interaction functions are defined only by some properties determining their shapes. To account for undercrowding phenomena, the prey growth function is assumed to model a strong Allee effect; to simulate the predator interference during the predation process, the trophic function is assumed predator-dependent. A stability analysis of the system is performed, using the predation efficiency and a measure of the predator interference as bifurcation parameters. The admissible scenarios are much richer than in the case of prey-dependent trophic functions, investigated in Buffoni et al. (2011). General conditions for the number of equilibria, for the existence and stability of extinction and coexistence equilibrium states are determined, and the bifurcations exhibited by the system are investigated. Numerical results illustrate the qualitative behaviours of the system, in particular the presence of limit cycles, of global bifurcations and of bistability situations.     

A numerical study of the formation of spatial patterns in twospotted spider mites

The aim of this paper is to study the formation of spatial patterns in a predator–prey system with Tetranychus urticae as prey and Phytoseiulus persimilis as predator. Logistic Lotka–Volterra predator–prey equations are solved numerically with two different response functions, two initial conditions and one data set. The spatial patterns are generated by introducing diffusion-driven instability in the predator–prey system. Among all parameters involved in predator–prey equations, only the predator interference parameter is varied to generate diffusion-driven instability leading to spatial patterns of population density. Spatial patterns are further generated with the inclusion of prey-taxis in the predator–prey system. Routh–Hurwitz’s conditions for stability are used to create instability with prey-taxis in the system. It is shown that it is possible to generate spatial patterns with zero flux boundary conditions even in a smaller domain with a suitable value of the predator interference parameter or prey-taxis.     

Adaptive control of nonlinear complex Holling II predator–prey system with unknown parameters

In recent years, control of nonlinear complex predator–prey systems has attracted the attention of many researchers. The previous works have some weaknesses such as neglecting the consideration of the effects of both model uncertainties and unknown parameters and having an infinite time of convergence. To overcome the mentioned shortages, this article solves the problem of robust control of nonlinear complex Holling type II predator–prey system in a given finite time. It is assumed that the parameters of the system are fully unknown in advance and some uncertainties perturb the system's dynamics. To tackle the system unknown parameters, some adaptation laws are introduced. Thereafter, a robust switching controller is proposed to finite‐timely stabilize the predator–prey system. An illustrative example demonstrates the efficiency and usefulness of the proposed control strategy. © 2015 Wiley Periodicals, Inc. Complexity 21: 260–266, 2016     

非自治Lotka-Volterra扩散模型的持续生存与周期轨道(英)

本文研究了一类非自治的捕食者一食饵扩散模型；其中食饵能在环境相异的两个缀块间有限制地扩散，但对捕食者来说，缀块间的扩散不受任何限制；另外假设模型的系数都是时间的函数．我们证明了在适当的条件下，这个系统能够持续生存，进一步给出了系统存在唯一全局渐近稳定正周期轨道的充分条件．     

A MICROFOUNDATION OF PREDATOR‐PREY DYNAMICS

ABSTRACT. Predator‐prey relationships account for an important part of all interactions betweenspecies. In this paper we provide a microfoundation for such predator‐prey relations in afood chain. Basic entities of our analysis are representative organisms of species modeled similar to economic households. With prices as indicators of scarcity, organisms are assumed to behave as if they maximize their net biomass subject to constraints which express the organisms' risk of being preyed upon during predation. Like consumers, organisms face a ‘budget constraint’ requiring their expenditure on prey biomass not to exceed their revenue from supplying own biomass. Short‐run ecosystem equilibria are defined and derived. The net biomass acquired by the representative organism in the short term determines the positive or negative population growth. Moving short‐run equilibria constitute the dynamics of the predator‐prey relations that are characterized in numerical analysis. The population dynamics derived here turn out to differ significantly from those assumed in the standard Lotka‐Volterra model.     

The dynamical behavior of a predator–prey system with Gompertz growth function and impulsive dispersal of prey between two patches

In this paper, we investigate a predator–prey model with Gompertz growth function and impulsive dispersal of prey between two patches. Using the dynamical properties of single‐species model with impulsive dispersal in two patches and comparison principle of impulsive differential equations, necessary and sufficient criteria on global attractivity of predator‐extinction periodic solution and permanence are established. Finally, a numerical example is given to illustrate the theoretical results. Copyright © 2015 John Wiley & Sons, Ltd.     

Permanence for a delayed periodic predator-prey model with prey dispersal in multi-patches and predator density-independent

In this paper, we study two species time-delayed predator-prey Lotka-Volterra type dispersal systems with periodic coefficients, in which the prey species can disperse among n patches, while the density-independent predator species is confined to one of patches and cannot disperse. Sufficient conditions on the boundedness, permanence and existence of positive periodic solution for this systems are established. The theoretical results are confirmed by a special example and numerical simulations.     

Food chain model with optimal harvesting in fuzzy environment

A multispecies harvesting model with mutual interactions is formulated based on Lotka–Voltera model with three competing species which are affected not only by harvesting but also by the presence of prey, predator and the third species, which is super predator. In order to understand the dynamics of the system, it is assumed that the super predator follows the logistic growth. Further, there is demand for all the above three species in the market and hence harvesting of all species is performed. We derive the condition for global stability of the system using a suitable Lyapunov function. The possibility of existence of bioeconomic equilibrium is discussed. The optimal harvest policy is studied and the solution is derived under imprecise inflation in fuzzy environment using Pontryagin’s maximal principle. Finally some numerical examples are discussed to illustrate the model.     

Permanence and periodicity of a delayed ratio-dependent predator–prey model with Holling type functional response and stage structure

A periodic and delayed ratio-dependent predator–prey system with Holling type III functional response and stage structure for both prey and predator is investigated. It is assumed that immature predator and mature individuals of each species are divided by a fixed age, and immature predator do not have the ability to attack prey. Sufficient conditions are derived for the permanence and existence of positive periodic solution of the model. Numerical simulations are presented to illustrate the feasibility of our main results.     

A MICRO‐LEVEL ‘CONSUMER APPROACH’ TO SPECIES POPULATION DYNAMICS

ABSTRACT. In this paper we develop a micro ecosystem model whose basic entities are representative organisms which behave as if maximizing their net offspring under constraints. Net offspring is increasing in prey biomass intake, declining in the loss of own biomass to predators and Allee's law applies. The organism's constraint reflects its perception of how scarce its own biomass and the biomass of its prey is. In the short‐run periods prices (scarcity indicators) coordinate and determine all biomass transactions and net offspring which directly translates into population growth functions. We are able to explicitly determine these growth functions for a simple food web when specific parametric net offspring functions are chosen in the micro‐level ecosystem model. For the case of a single species our model is shown to yield the well‐known Verhulst‐Pearl logistic growth function. With two species in predator‐prey relationship, we derive differential equations whose dynamics are completely characterized and turn out to be similar to the predator‐prey model with Michaelis‐Menten type functional response. With two species competing for a single resource we find that coexistence is a knife‐edge feature confirming Tschirhart's [2002] result in a different but related model.     

Uniqueness and stability of positive steady state solutions for a ratio-dependent predator–prey system with a crowding term in the prey equation

This paper deals with a ratio-dependent predator–prey system with a crowding term in the prey equation, where it is assumed that the coefficient of the functional response is less than the coefficient of the intrinsic growth rates of the prey species. We demonstrate some special behaviors of solutions to the system which the coexistence states of two species can be obtained when the crowding region in the prey equation only is designed suitably. Furthermore, we demonstrate that under some conditions, the positive steady state solution of the predator–prey system with a crowding term in the prey equation is unique and stable. Our result is different from those ones of the predator–prey systems without the crowding terms.     

PERSISTENCE AND GLOBAL STABILITY FOR TWO-SPECIES NONAUTONOMOUS PREDATOR-PREY SYSTEM WITH DIFFUSION AND TIME DELAY

In this paper, a non-autonomous predator-prey model with diffusion andcontinuous time delay is studied, where the prey can diffuse between two pat-ches of a heterogeneous environment with barriers between patches, but for thepredator, the diffusion does not involve a barrier between patches, further itis assumed that all the parameters are time-dependent. It is shown that thesystem can be made persistent under some appropriate conditions. Moreover,sufficient conditions that guarantee the existence of a unique periodic solutionwhich is globally asymptotic stable are derived.     

Permanence and extinction analysis for a delayed periodic predator-prey system with Holling type II response function and diffusion

In this paper, it is studied that two species predator-prey Lotka-Volterra type dispersal system with delay and Holling type II response function, in which the prey species can disperse among n patches, while the density-independent predator species is confined to one of the patches and cannot disperse. Sufficient conditions of integrable form for the boundedness, permanence, extinction and the existence of positive periodic solution are established, respectively.