Turing patterns of a strongly coupled predator-prey system with diffusion effects

The main purpose of this work is to investigate the effects of cross-diffusion in a strongly coupled predator-prey system. By a linear stability analysis we find the conditions which allow a homogeneous steady state (stable for the kinetics) to become unstable through a Turing mechanism. In particular, it is shown that Turing instability of the reaction-diffusion system can disappear due to the presence of the cross-diffusion, which implies that the cross-diffusion induced stability can be regarded as the cross-stability of the corresponding reaction-diffusion system. Furthermore, we consider the existence and non-existence results concerning non-constant positive steady states (patterns) of the system. We demonstrate that cross-diffusion can create non-constant positive steady-state solutions. These results exhibit interesting and very different roles of the cross-diffusion in the formation and the disappearance of the Turing instability.     

A weighted quasilinear equation related to the mean curvature operator

This paper studies a weighted quasilinear perturbation, through the mean curvature flow operator, of the classical linear heat equation. The mean curvature has the effect of maintaining bounded all classical positive steady states of the model, though their derivatives must be somewhere unbounded. The dynamics of the positive solutions of the model is fully described.     

Qualitative analysis of steady states to the Sel'kov model

In this work, we are concerned with a reaction-diffusion system well known as the Sel'kov model, which has been used for the study of morphogenesis, population dynamics and autocatalytic oxidation reactions. We derive some further analytic results for the steady states to this model. In particular, we show that no nonconstant positive steady state exists if 0<p?1 and θ is large, which provides a sharp contrast to the case of p>1 and large θ, where nonconstant positive steady states can occur. Thus, these conclusions indicate that the parameter p plays a crucial role in leading to spatially nonhomogeneous distribution of the two reactants. The a priori estimates are fundamental to our mathematical approaches.     

Travelling wave fronts in reaction-diffusion systems with spatio-temporal delays

This paper deals with the existence of travelling wave fronts in reaction-diffusion systems with spatio-temporal delays. Our approach is to use monotone iterations and a nonstandard ordering for the set of profiles of the corresponding wave system. New iterative techniques are established for a class of integral operators when the reaction term satisfies different monotonicity conditions. Following this, the existence of travelling wave fronts for reaction-diffusion systems with spatio-temporal delays is established. Finally, we apply the main results to a single-species diffusive model with spatio-temporal delay and obtain some existence criteria of travelling wave fronts by choosing different kernels.     

偏微分方程的吸引子

Abstract. In this paper the reaction-diffusion systems and the damped wave systems with non-linear terms of gradient form are studied ,and the conditions for making them to be gradient sys-tems are given. So the structure of attractors is clear and simple in some sense by the known re-sult. Some examples are given. Also the reaction-diffusion systems with general nonlinear termsare discussed.     

Hair-triggered instability of radial steady states, spread and extinction in semilinear heat equations

We first study the initial value problem for a general semilinear heat equation. We prove that every bounded nonconstant radial steady state is unstable if the spatial dimension is low (n?10) or if the steady state is flat enough at infinity: the solution of the heat equation either becomes unbounded as t approaches the lifespan, or eventually stays above or below another bounded radial steady state, depending on if the initial value is above or below the first steady state; moreover, the second steady state must be a constant if n?10.Using this instability result, we then prove that every nonconstant radial steady state of the generalized Fisher equation is a hair-trigger for two kinds of dynamical behavior: extinction and spreading. We also prove more criteria on initial values for these types of behavior. Similar results for a reaction-diffusion system modeling an isothermal autocatalytic chemical reaction are also obtained.     

Pattern formation in a nonlinear model for animal coats

Several models have been proposed for describing the formation of animal coat patterns. We consider reaction-diffusion models due to Murray, which rely on a Turing instability for the pattern selection. In this paper, we describe the early stages of the pattern formation process for large domain sizes. This includes the selection mechanism and the geometry of the patterns generated by the nonlinear system on one-, two-, and three-dimensional base domains. These results are obtained by an adaptation of results explaining the occurrence of spinodal decomposition in materials science as modeled by the Cahn-Hilliard equation. We use techniques of dynamical systems, viewing solutions of the reaction-diffusion model in terms of nonlinear semiflows. Our results are applicable to any parabolic system exhibiting a Turing instability.     

The instability of spiky steady states for a competing species model with cross diffusion

This paper is concerned with the stability/instability of a class of positive spiky steady states for a quasi-linear cross-diffusion system describing two-species competition. By detailed spectral analysis, it is proved that the spiky steady states for the related shadow system are linearly unstable and the spiky steady states for the original cross-diffusion system are non-linearly unstable.     

Uniqueness and asymptotic behavior of coexistence states for a non-cooperative model of nuclear reactors

In this paper, we analyze the asymptotic behavior of coexistence states for a non-cooperative model of nuclear reactors. In addition, we also present some remarks on the uniqueness of coexistence states in a high dimensional case. Our results complement the work of López-Gómez [J. López-Gómez, The steady states of a non-cooperative model of nuclear reactors, J. Differential Equations 246 (2009), 358-372].     

Turing pattern of the Oregonator model

The Oregonator model is the mathematical dynamics which describes the Field-Körös-Noyes mechanics of the famous Belousov-Zhabotinskii? reaction. In this work, we establish some fundamental analytic properties of this dynamics and its corresponding steady state. Under various conditions on the parameters and the size of the reactor, we examine the existence and non-existence of non-constant steady states. In particular, for some properly chosen parameter ranges, we prove the occurrence of the Turing pattern generated by this Oregonator model. Our results exhibit interesting and very different roles of the diffusion rates and the reactor in the formation of the Turing pattern. Our mathematical analysis mainly relies on a priori estimates and the topological degree argument.     

Non-constant positive steady states of the Sel'kov model

This paper deals with the reaction-diffusion system known as the Sel'kov model with the homogeneous Neumann boundary condition. This model has been applied to various problems in chemistry and biology. We first give a priori estimates (positive upper and lower bounds) of positive steady states, and then study the non-existence, bifurcation and global existence of non-constant positive steady states as the parameters λ and θ are varied.     

Traveling wave solutions for reaction-diffusion systems

This paper is concerned with traveling waves of reaction-diffusion systems. The definition of coupled quasi-upper and quasi-lower solutions is introduced for systems with mixed quasimonotone functions, and the definition of ordered quasi-upper and quasi-lower solutions is also given for systems with quasimonotone nondecreasing functions. By the monotone iteration method, it is shown that if the system has a pair of coupled quasi-upper and quasi-lower solutions, then there exists at least a traveling wave solution. Moreover, if the system has a pair of ordered quasi-upper and quasi-lower solutions, then there exists at least a traveling wavefront. As an application we consider the delayed system of a mutualistic model.     

Perron Theorem in the monotone iteration method for traveling waves in delayed reaction-diffusion equations

In this paper we revisit the existence of traveling waves for delayed reaction-diffusion equations by the monotone iteration method. We show that Perron Theorem on existence of bounded solution provides a rigorous and constructive framework to find traveling wave solutions of reaction-diffusion systems with time delay. The method is tried out on two classical examples with delay: the predator-prey and Belousov-Zhabotinskii models.     

Hopf bifurcations in a reaction-diffusion population model with delay effect

A reaction-diffusion population model with a general time-delayed growth rate per capita is considered. The growth rate per capita can be logistic or weak Allee effect type. From a careful analysis of the characteristic equation, the stability of the positive steady state solution and the existence of forward Hopf bifurcation from the positive steady state solution are obtained via the implicit function theorem, where the time delay is used as the bifurcation parameter. The general results are applied to a “food-limited” population model with diffusion and delay effects as well as a weak Allee effect population model.     

Diffusive monotonicity and threshold dynamics of delayed reaction diffusion equations

For abstract functional differential equations and reaction-diffusion equations with delay, an exponential ordering is introduced which takes into account the spatial diffusion. The induced monotonicity of the solution semiflows is established and applied to describe the threshold dynamics (extinction or persistence/convergence to positive equilibria) for a nonlocal and delayed reaction-diffusion population model.     

Asymptotic profiles of the positive steady state for an SIS epidemic reaction-diffusion model. Part I

To capture the impact of spatial heterogeneity of environment and movement of individuals on the persistence and extinction of a disease, Allen et al. in [L.J.S. Allen, B.M. Bolker, Y. Lou, A.L. Nevai, Asymptotic profiles of the steady states for an SIS epidemic reaction-diffusion model, Discrete Contin. Dyn. Syst. Ser. A 21 (1) (2008) 1-20] proposed a spatial SIS (susceptible-infected-susceptible) reaction-diffusion model, and studied the existence, uniqueness and particularly the asymptotic behavior of the endemic equilibrium as the diffusion rate of the susceptible individuals goes to zero in the case where a so-called low-risk subhabitat is created. In this work, we shall provide further understanding of the impacts of large and small diffusion rates of the susceptible and infected population on the persistence and extinction of the disease, which leads us to determine the asymptotic behaviors of the endemic equilibrium when the diffusion rate of either the susceptible or infected population approaches to infinity or zero in the remaining cases. Consequently, our results reveal that, in order to eliminate the infected population at least in low-risk area, it is necessary that one will have to create a low-risk subhabitat and reduce at least one of the diffusion rates to zero. In this case, our results also show that different strategies of controlling the diffusion rates of individuals may lead to very different spatial distributions of the population; moreover, once the spatial environment is modified to include a low-risk subhabitat, the optimal strategy of eradicating the epidemic disease is to restrict the diffusion rate of the susceptible individuals rather than that of the infected ones.     

Non-existence of non-constant positive steady states of two Holling type-II predator-prey systems: Strong interaction case

We prove the non-existence of non-constant positive steady state solutions of two reaction-diffusion predator-prey models with Holling type-II functional response when the interaction between the predator and the prey is strong. The result implies that the global bifurcating branches of steady state solutions are bounded loops.     

Spectral properties of non-local differential operators

In this article we consider the spectral properties of a class of non-local operators that arise from the study of non-local reaction-diffusion equations. Such equations are used to model a variety of physical and biological systems with examples ranging from Ohmic heating to population dynamics. The operators studied here are bounded perturbations of linear (local) differential operators. The non-local perturbation is in the form of an integral term. It is shown here that the spectral properties of these non-local operators can differ considerably from those of their local counterpart. Multiplicities of eigenvalues are studied and new oscillation results for the associated eigenfunctions are presented. These results highlight problems with certain similar results and provide an alternative formulation. Finally, the stability of steady states of associated non-local reaction-diffusion equations is discussed.     

Persistence of periodic patterns for perturbed biological oscillators

We apply the perturbation method and the Implicit Function Theorem to study the persistence of periodic patterns for two perturbed general biological oscillators: one weakly coupled ordinary differential equation and one reaction-diffusion system. The proof relies also on the formal adjoint equation theory and the Lyapunov-Schmidt method. For the perturbed reaction-diffusion model, spatial-temporal patterns such as rotating waves are shown to exist. Numerical simulations are presented to illustrate our analytical results.     

Persistence of wavefronts in delayed nonlocal reaction-diffusion equations

We develop a perturbation argument based on existing results on asymptotic autonomous systems and the Fredholm alternative theory that yields the persistence of traveling wavefronts for reaction-diffusion equations with nonlocal and delayed nonlinearities, when the time lag is relatively small. This persistence result holds when the nonlinearity of the corresponding ordinary reaction-diffusion system is either monostable or bistable. We then illustrate this general result using five different models from population biology, epidemiology and bio-reactors.