Two-parameter bifurcations in a network of two neurons with multiple delays

We consider a network of two coupled neurons with delayed feedback. We show that the connection topology of the network plays a fundamental role in classifying the rich dynamics and bifurcation phenomena. Regarding eigenvalues of the connection matrix as bifurcation parameters, we obtain codimension 1 bifurcations (including a fold bifurcation and a Hopf bifurcation) and codimension 2 bifurcations (including fold-Hopf bifurcations and Hopf-Hopf bifurcations). We also give concrete formulae for the normal form coefficients derived via the center manifold reduction that give detailed information about the bifurcation and stability of various bifurcated solutions. In particular, we obtain stable or unstable equilibria, periodic solutions, quasi-periodic solutions, and sphere-like surfaces of solutions. We also show how to evaluate critical normal form coefficients from the original system of delay-differential equations without computing the corresponding center manifolds.     

Dynamics of two-cell systems with discrete delays

We consider the system of delay differential equations (DDE) representing the models containing two cells with time-delayed connections. We investigate global, local stability and the bifurcations of the trivial solution under some generic conditions on the Taylor coefficients of the DDE. Regarding eigenvalues of the connection matrix as bifurcation parameters, we obtain codimension one bifurcations (including pitchfork, transcritical and Hopf bifurcation) and Takens-Bogdanov bifurcation as a codimension two bifurcation. For application purposes, this is important since one can now identify the possible asymptotic dynamics of the DDE near the bifurcation points by computing quantities which depend explicitly on the Taylor coefficients of the original DDE. Finally, we show that the analytical results agree with numerical simulations.     

Bifurcation analysis on a generalized recurrent neural network with two interconnected three-neuron components

A class of recurrent neural networks is constructed by generalizing a specific class of n-neuron networks. It is shown that the newly constructed network experiences generic pitchfork and Hopf codimension one bifurcations. It is also proved that the emergence of generic Bogdanov–Takens, pitchfork–Hopf and Hopf–Hopf codimension two, and the degenerate Bogdanov–Takens bifurcation points in the parameter space is possible due to the intersections of codimension one bifurcation curves. The occurrence of bifurcations of higher codimensions significantly increases the capability of the newly constructed recurrent neural network to learn broader families of periodic signals.     

Versal unfoldings of predator-prey systems with ratio-dependent functional response

In this paper we study the versal unfolding of a predator-prey system with ratio-dependent functional response near a degenerate equilibrium in order to obtain all possible phase portraits for its perturbations. We first construct the unfolding and prove its versality and degeneracy of codimension 2. Then we discuss all its possible bifurcations, including transcritical bifurcation, Hopf bifurcation, and heteroclinic bifurcation, give conditions of parameters for the appearance of closed orbits and heteroclinic loops, and describe the bifurcation curves. Phase portraits for all possible cases are presented.     

Principal Asymptotics in the Problem on the Andronov–Hopf Bifurcation and Their Applications

New formulas are obtained for the principal asymptotics of bifurcation solutions in the problem on the Andronov–Hopf bifurcation, leading to new algorithms for studying bifurcations in the general setting. The approach proposed in the paper allows one to consider not only the classical problems about bifurcations of codimension one but also some problems concerning bifurcations of codimension two. A new approach to the analysis of bifurcations of cycles in systems with homogeneous nonlinearities is proposed. As an application, we consider the problem on the bifurcation of periodic solutions of the van der Pol equation.     

Bifurcation from periodic solutions with spatiotemporal symmetry, including resonances and mode interactions

We study local bifurcation in equivariant dynamical systems from periodic solutions with a mixture of spatial and spatiotemporal symmetries.In previous work, we focused primarily on codimension one bifurcations. In this paper, we show that the techniques used in the codimension one analysis can be extended to understand also higher codimension bifurcations, including resonant bifurcations and mode interactions. In particular, we present a general reduction scheme by which we relate bifurcations from periodic solutions to bifurcations from fixed points of twisted equivariant diffeomorphisms, which in turn are linked via normal form theory to bifurcations from equilibria of equivariant vector fields.We also obtain a general theory for bifurcation from relative periodic solutions and we show how to incorporate time-reversal symmetries into our framework.     

Bifurcation analysis of a generalized Gause model with prey harvesting and a generalized Holling response function of type III

In this paper we study a generalized Gause model with prey harvesting and a generalized Holling response function of type III: . The goal of our study is to give the bifurcation diagram of the model. For this we need to study saddle-node bifurcations, Hopf bifurcation of codimension 1 and 2, heteroclinic bifurcation, and nilpotent saddle bifurcation of codimension 2 and 3. The nilpotent saddle of codimension 3 is the organizing center for the bifurcation diagram. The Hopf bifurcation is studied by means of a generalized Liénard system, and for b=0 we discuss the potential integrability of the system. The nilpotent point of multiplicity 3 occurs with an invariant line and can have a codimension up to 4. But because it occurs with an invariant line, the effective highest codimension is 3. We develop normal forms (in which the invariant line is preserved) for studying of the nilpotent saddle bifurcation. For b=0, the reversibility of the nilpotent saddle is discussed. We study the type of the heteroclinic loop and its cyclicity. The phase portraits of the bifurcations diagram (partially conjectured via the results obtained) allow us to give a biological interpretation of the behavior of the two species.     

Anti-control of Hopf bifurcation in the new chaotic system with two stable node-foci

In order to further understand a complex 3D dynamical system showing strange chaotic attractors with two stable node-foci near Hopf bifurcation point, we propose nonlinear control scheme to the system and the controlled system, depending on five parameters, can exhibit codimension one, two, and three Hopf bifurcations in a much larger parameter regain. The control strategy used keeps the equilibrium structure of the chaotic system and can be applied to degenerate Hopf bifurcation at the desired location with preferred stability.     

Limit cycles in slow-fast codimension 3 saddle and elliptic bifurcations

This paper deals with local bifurcations occurring near singular points of planar slow-fast systems. In particular, it is concerned with the study of the slow-fast variant of the unfolding of a codimension 3 nilpotent singularity. The slow-fast variant of a codimension 1 Hopf bifurcation has been studied extensively before and its study has lead to the notion of canard cycles in the Van der Pol system. Similarly, codimension 2 slow-fast Bogdanov–Takens bifurcations have been characterized. Here, the singularity is of codimension 3 and we distinguish slow-fast elliptic and slow-fast saddle bifurcations. We focus our study on the appearance on small-amplitude limit cycles, and rely on techniques from geometric singular perturbation theory and blow-up.     

Analyses of Bifurcations and Stability in a Predator-prey System with Holling Type-Ⅳ Functional Response

In this paper the dynamical behaviors of a predator-prey system with Holling Type-Ⅳfunctionalresponse are investigated in detail by using the analyses of qualitative method,bifurcation theory,and numericalsimulation.The qualitative analyses and numerical simulation for the model indicate that it has a unique stablelimit cycle.The bifurcation analyses of the system exhibit static and dynamical bifurcations including saddle-node bifurcation,Hopf bifurcation,homoclinic bifurcation and bifurcation of cusp-type with codimension two(ie,the Bogdanov-Takens bifurcation),and we show the existence of codimension three degenerated equilibriumand the existence of homoclinic orbit by using numerical simulation.     

Codimension two bifurcations of fixed points in a class of vibratory systems with symmetrical rigid stops

A vibratory system having symmetrically placed rigid stops and subjected to periodic excitation is considered. Local codimension two bifurcations of the vibratory system with symmetrical rigid stops, associated with double Hopf bifurcation and interaction of Hopf and pitchfork bifurcation, are analyzed by using the center manifold theorem technique and normal form method of maps. Dynamic behavior of the system, near the points of codimension two bifurcations, is investigated by using qualitative analysis and numerical simulation. Hopf-flip bifurcation of fixed points in the vibratory system with a single stop are briefly analyzed by comparison with unfoldings analyses of Hopf-pitchfork bifurcation of the vibratory system with symmetrical rigid stops. Near the value of double Hopf bifurcation there exist period-one double-impact symmetrical motion and quasi-periodic impact motions. The quasi-periodic impact motions are represented by the closed circle and “tire-like” attractor in projected Poincaré sections. With change of system parameters, the quasi-periodic impact motions usually lead to chaos via “tire-like” torus doubling.     

Symmetric functional differential equations and neural networks with memory

We establish an analytic local Hopf bifurcation theorem and a topological global Hopf bifurcation theorem to detect the existence and to describe the spatial-temporal pattern, the asymptotic form and the global continuation of bifurcations of periodic wave solutions for functional differential equations in the presence of symmetry. We apply these general results to obtain the coexistence of multiple large-amplitude wave solutions for the delayed Hopfield-Cohen-Grossberg model of neural networks with a symmetric circulant connection matrix.

     

Bifurcation analysis of a Leslie-type predator–prey system with simplified Holling type IV functional response and strong Allee effect on prey

In this paper, a Leslie-type predator–prey system with simplified Holling type IV functional response and strong Allee effect on prey is proposed. The dissipativity of the system and the existence of all possible equilibria are investigated. The investigation emphasizes the exploring of bifurcation. It is shown that the system exists several non-hyperbolic positive equilibria, such as a weak focus of multiplicities one and two, (degenerate) saddle–nodes and Bogdanov–Takens singularities (cusp case) of codimensions 2 and 3. At these equilibria, it is proved that the system undergoes various kinds of bifurcations, such as saddle–node bifurcation, Hopf bifurcation, degenerate Hopf bifurcation and Bogdanov–Takens bifurcation of codimensions 2 and 3. With the parameters selected properly, there exhibits a limit cycle, a homoclinic loop, two limit cycles, a semistable limit cycle, or the simultaneous occurrence of a homoclinic loop and a limit cycle in the system. Moreover, it is also proved that the system has a cusp of codimension at least 4. Hence, there may exist three limit cycles generated from Hopf bifurcation of codimension 3. Numerical simulations are done to support the theoretical results.     

Bogdanov‐Takens bifurcations of codimensions 2 and 3 in a Leslie‐Gower predator‐prey model with Michaelis‐Menten–type prey harvesting

The Bogdanov‐Takens bifurcations of a Leslie‐Gower predator‐prey model with Michaelis‐Menten–type prey harvesting were studied. In the paper “Diff. Equ. Dyn. Syst. 20(2012), 339‐366,” Gupta et al proved that the Leslie‐Gower predator‐prey model with Michaelis‐Menten–type prey harvesting has rich dynamics. Some equilibria of codimension 1 and their bifurcations were discussed. In this paper, we find that the model has an equilibrium of codimensions 2 and 3. We also prove analytically that the model undergoes Bogdanov‐Takens bifurcations (cusp cases) of codimensions 2 and 3. Hence, the model can have 2 limit cycles, coexistence of a stable homoclinic loop and an unstable limit cycle, supercritical and subcritical Hopf bifurcations, and homoclinic bifurcation of codimension 1 as the values of parameters vary. Moreover, several numerical simulations are conducted to illustrate the validity of our results.     

Bifurcation analysis on a survival red blood cells model

In this paper, we consider a model described the survival of red blood cells in animal. Its dynamics are studied in terms of local and global Hopf bifurcations. We show that a sequence of Hopf bifurcations occur at the positive equilibrium as the delay crosses some critical values. Using the reduced system on the center manifold, we also obtain that the periodic orbits bifurcating from the positive equilibrium are stable in the center manifold, and all Hopf bifurcations are supercritical. Further, particular attention is focused on the continuation of local Hopf bifurcation. We show that global Hopf bifurcations exist after the second critical value of time delay.     

Hopf‐pitchfork bifurcation in a two‐neuron system with discrete and distributed delays

Both discrete and distributed delays are considered in a two‐neuron system. We analyze the influence of interaction coefficient and time delay on the Hopf‐pitchfork bifurcation. First, we obtain the codimension‐2 unfolding with original parameters for Hopf‐pitchfork bifurcation by using the center manifold reduction and the normal form method. Next, through analyzing the unfolding structure, we give complete bifurcation diagrams and phase portraits, in which multistability and other dynamical behaviors of the original system are found, such as a stable periodic orbit, the coexistence of two stable nontrivial equilibria, and the coexistence of a stable periodic orbit and two stable equilibria. In addition, the obtained theoretical results are verified by numerical simulations. Finally, we perform the comparisons of the obtained results of Hopf‐pitchfork bifurcation with other Hopf‐fold bifurcation results in some biological neural systems and give the obtained mathematical results corresponding to the physical states of neurons. Copyright © 2015 JohnWiley & Sons, Ltd.     

Complex Dynamics in a Unified SIR and HIV Disease Model: A Bifurcation Theory Approach

This paper is concerned with complex dynamical behaviors of a simple unified SIR and HIV disease model with a convex incidence and four real parameters. Due to the complex nature of the disease dynamics, our goal is to explore bifurcations including multistable states, limit cycles, and homoclinic loops in the whole parameter space. The first contribution is the proof of the existence of multiple limit cycles giving rise from Hopf bifurcation, which further induces bistable or tristable states because of the coexistence of stable equilibria and periodic motion. Next, we propose that the existence of Bogdanov–Takens (BT) bifurcation yields the bifurcation of homoclinic loops, which provides a new mechanism for generating disease recurrence, for example, the relapse–remission, viral blip cycles in HIV infection. Last, we present a novel method for the derivation of the normal forms of codimension two and three BT bifurcations. The method is based on the simplest normal form theory from Yu’s previous works.

     

Synchronization for two coupled oscillators with inhibitory connection

In this paper we present an oscillatory neural network composed of two coupled neural oscillators with inhibitory connections. Each of the oscillators describes the dynamics of average activities of excitatory and inhibitory populations of neurons. Regarding time delays τ as the bifurcation parameter, we not only obtain the existence of Hopf bifurcations but also investigate the bifurcation direction and stability of bifurcated periodic solutions by employing normal form theory and center manifold reduction. Finally, numerical simulations are provided to illustrate the theoretical results. Copyright © 2009 John Wiley & Sons, Ltd.     

Bifurcations of a Leslie-Gower prey-predator system with ratio-dependent Holling IV functional response and prey harvesting

The dynamics of a Leslie-Gower prey-predator system with ratio-dependent Holling IV functional response and constant harvesting rate of prey are taken into account. The results developed in this article reveal far richer dynamics compared with the system without harvesting. We first make qualitative and bifurcation analysis of the system without harvesting and show that the system has a weak focus of multiplicity at most 2, at which a Hopf bifurcation occurs. However, the system with harvesting has four nonhyperbolic equilibria for some parameter values, such as two saddle-node, a cusp, and a weak focus of multiplicity at most 4, and exhibits two saddle-node bifurcations, a Bogdanov-Takens bifurcation of codimension 2, and a Hopf bifurcation. It reveals that there exist some critical harvesting values such that the species are in danger of extinction when the harvesting rate is greater than the critical values, which indicates that the dynamics of the system are sensitive to the constant prey harvesting. Moreover, numerical simulations are presented to illustrate our theoretical results.     

Global Hopf bifurcation in the Leslie-Gower predator-prey model with two delays

In this paper, the Leslie-Gower predator-prey system with two delays is investigated. By choosing the delay as a bifurcation parameter, we show that Hopf bifurcations can occur as the delay crosses some critical values. In addition, special attention is paid to the global continuation of local Hopf bifurcations. Using a global Hopf bifurcation theorem for functional differential equations, we show the global existence of periodic solutions.