Effect of emergent carrying capacity in an eco‐epidemiological system

The paper explores an eco‐epidemiological model of a predator–prey type, where the prey population is subject to infection. The model is basically a combination of S‐I type model and a Rosenzweig–MacArthur predator–prey model. The novelty of this contribution is to consider different competition coefficients within the prey population, which leads to the emergent carrying capacity. We explicitly separate the competition between non‐infected and infected individuals. This emergent carrying capacity is markedly different to the explicit carrying capacities that have been considered in many eco‐epidemiological models. We observed that different intra‐class and inter‐class competition can facilitate the coexistence of susceptible prey‐infected prey–predator, which is impossible for the case of the explicit carrying capacity model. We also show that these findings are closely associated with bi‐stability. The present system undergoes bi‐stability in two different scenarios: (a) bi‐stability between the planner equilibria where susceptible prey co‐exists with predator or infected prey and (b) bi‐stability between co‐existence equilibrium and the planner equilibrium where susceptible prey coexists with infected prey; have been discussed. The conditions for which the system is to be permanent and the global stability of the system around disease‐free equilibrium are worked out. Copyright © 2015 John Wiley & Sons, Ltd.     

Dynamics and pattern formations in diffusive predator‐prey models with two prey‐taxis

A reaction‐diffusion two‐predator‐one‐prey system with prey‐taxis describes the spatial interaction and random movement of predator and prey species, as well as the spatial movement of predators pursuing preys. The global existence and boundedness of solutions of the system in bounded domains of arbitrary spatial dimension and any small prey‐taxis sensitivity coefficient are investigated by the semigroup theory. The spatial pattern formation induced by the prey‐taxis is characterized by the Turing type linear instability of homogeneous state; it is shown that prey‐taxis can both compress and prompt the spatial patterns produced through diffusion‐induced instability in two‐predator‐one‐prey systems.     

A MICRO‐LEVEL ‘CONSUMER APPROACH’ TO SPECIES POPULATION DYNAMICS

ABSTRACT. In this paper we develop a micro ecosystem model whose basic entities are representative organisms which behave as if maximizing their net offspring under constraints. Net offspring is increasing in prey biomass intake, declining in the loss of own biomass to predators and Allee's law applies. The organism's constraint reflects its perception of how scarce its own biomass and the biomass of its prey is. In the short‐run periods prices (scarcity indicators) coordinate and determine all biomass transactions and net offspring which directly translates into population growth functions. We are able to explicitly determine these growth functions for a simple food web when specific parametric net offspring functions are chosen in the micro‐level ecosystem model. For the case of a single species our model is shown to yield the well‐known Verhulst‐Pearl logistic growth function. With two species in predator‐prey relationship, we derive differential equations whose dynamics are completely characterized and turn out to be similar to the predator‐prey model with Michaelis‐Menten type functional response. With two species competing for a single resource we find that coexistence is a knife‐edge feature confirming Tschirhart's [2002] result in a different but related model.     

A MICROFOUNDATION OF PREDATOR‐PREY DYNAMICS

ABSTRACT. Predator‐prey relationships account for an important part of all interactions betweenspecies. In this paper we provide a microfoundation for such predator‐prey relations in afood chain. Basic entities of our analysis are representative organisms of species modeled similar to economic households. With prices as indicators of scarcity, organisms are assumed to behave as if they maximize their net biomass subject to constraints which express the organisms' risk of being preyed upon during predation. Like consumers, organisms face a ‘budget constraint’ requiring their expenditure on prey biomass not to exceed their revenue from supplying own biomass. Short‐run ecosystem equilibria are defined and derived. The net biomass acquired by the representative organism in the short term determines the positive or negative population growth. Moving short‐run equilibria constitute the dynamics of the predator‐prey relations that are characterized in numerical analysis. The population dynamics derived here turn out to differ significantly from those assumed in the standard Lotka‐Volterra model.     

An ecoepidemiological predator‐prey model with standard disease incidence

This investigation accounts for epidemics spreading among interacting populations. The infective disease spreads among the prey, of which only susceptibles reproduce, while infected prey do not grow, recover, reproduce nor compete for resources. The model is general enough to describe a large number of ecosystems, on land, in the air or in the water. The main results concern the boundedness of the trajectories, the analysis of local and global stability, system's persistency and a threshold property below which the infection disappears. A sufficiently strong disease in the prey may avoid predators extinction and its presence can destabilize an otherwise stable predator‐prey configuration. The occurrence of transcritical, saddle‐node and Hopf‐bifurcations is also shown. Copyright © 2008 John Wiley & Sons, Ltd.     

Bogdanov‐Takens bifurcations of codimensions 2 and 3 in a Leslie‐Gower predator‐prey model with Michaelis‐Menten–type prey harvesting

The Bogdanov‐Takens bifurcations of a Leslie‐Gower predator‐prey model with Michaelis‐Menten–type prey harvesting were studied. In the paper “Diff. Equ. Dyn. Syst. 20(2012), 339‐366,” Gupta et al proved that the Leslie‐Gower predator‐prey model with Michaelis‐Menten–type prey harvesting has rich dynamics. Some equilibria of codimension 1 and their bifurcations were discussed. In this paper, we find that the model has an equilibrium of codimensions 2 and 3. We also prove analytically that the model undergoes Bogdanov‐Takens bifurcations (cusp cases) of codimensions 2 and 3. Hence, the model can have 2 limit cycles, coexistence of a stable homoclinic loop and an unstable limit cycle, supercritical and subcritical Hopf bifurcations, and homoclinic bifurcation of codimension 1 as the values of parameters vary. Moreover, several numerical simulations are conducted to illustrate the validity of our results.     

Hopf bifurcation of a diffusive Gause‐type predator‐prey model induced by time fractional‐order derivatives

Since population behaviors possess the characteristic of history memory, we, in this paper, introduce time fractional‐order derivatives into a diffusive Gause‐type predator‐prey model, which is time fractional‐order reaction‐diffusion equations and a generalized form of its corresponding first‐derivative model. For this kind of model, we prove the existence and uniqueness of a global positive solution by using the theory of evolution equations and the comparison principle of time fractional‐order partial differential equations. Besides, we obtain the stability and Hopf bifurcation of the Gause‐type predator‐prey model in the forms of the time fractional‐order ordinary equations and of the time fractional‐order reaction‐diffusion equations, respectively. Our results show that the stable region of the parameters in these 2 models can be enlarged by the time fractional‐order derivatives. Some numerical simulations are made to verify our results.     

Stability,Steady‐State Bifurcations,and Turing Patterns in a Predator–Prey Model with Herd Behavior and Prey‐taxis

In this paper, we consider a predator–prey model with herd behavior and prey‐taxis subject to the homogeneous Neumann boundary condition. First, by analyzing the characteristic equation, the local stability of the positive equilibrium is discussed. Then, choosing prey‐tactic sensitivity coefficient as the bifurcation parameter, we obtain a branch of nonconstant solutions bifurcating from the positive equilibrium by an abstract bifurcation theory, and find the stable bifurcating solutions near the bifurcation point under suitable conditions. We have shown that prey‐taxis can destabilize the uniform equilibrium and yields the occurrence of spatial patterns. Furthermore, some numerical simulations to illustrate the theoretical analysis are also carried out, Turing patterns such as spots pattern, spots–strip pattern, strip pattern, stable nonconstant steady‐state solutions, and spatially inhomogeneous periodic solutions are obtained, which also expand our theoretical results.     

Disease‐selective predation may lead to prey extinction

In real world bio‐communities, predational choice plays a key role to the persistence of the prey population. Predator's ‘sense’ of choice for predation towards the infected and noninfected prey is an important factor for those bio‐communities. There are examples where the predator can distinguish the infected prey and avoids those at the time of predation. Based on the examples, we propose two mathematical models and observe the dynamics of the systems around biologically feasible equilibria. For disease‐selective predation model there is a high risk of prey extinction. On the other hand, for non‐disease selective predation both populations co‐exist. Local stability analysis and global stability analysis of the positive interior equilibrium are performed. Moreover, conditions for the permanence of the system are obtained. Finally, we conclude that strictly disease‐selective predation may not be acceptable for the persistence of the prey population. Copyright © 2005 John Wiley & Sons, Ltd.     

The discrete‐time Kolmogorov systems with historic behavior

The main aim of this paper is to discuss some feature so‐called historic behavior of a discrete‐time Kolmogorov system of predator–prey interactions, which causes the nonexistence of the time averages.     

Existence and non‐existence of steady states to a cross‐diffusion system arising in a Leslie predator–prey model

This paper is concerned with a cross‐diffusion system arising in a Leslie predator–prey population model in a bounded domain with no flux boundary condition. We investigate sufficient condition for the existence and the non‐existence of non‐constant positive solution. We obtain that if natural diffusion coefficient of predator is large enough and cross‐diffusion coefficients are fixed, then under some conditions there exists non‐constant positive solution. Furthermore, we show that if natural diffusion coefficients of predator and prey are both large enough, and cross‐diffusion coefficients are small enough, then there exists no non‐constant positive solution. Copyright © 2012 John Wiley & Sons, Ltd.     

Dynamics of a ratio‐dependent stage‐structured predator‐prey model with delay

In this paper, we investigate the dynamics of a time‐delay ratio‐dependent predator‐prey model with stage structure for the predator. This predator‐prey system conforms to the realistically biological environment. The existence and stability of the positive equilibrium are thoroughly analyzed, and the sufficient and necessary conditions for the stability and instability of the positive equilibrium are obtained for the case without delay. Then, the influence of delay on the dynamics of the system is investigated using the geometric criterion developed by Beretta and Kuang. 26 We show that the positive steady state can be destabilized through a Hopf bifurcation and there exist stability switches under some conditions. The formulas determining the direction and the stability of Hopf bifurcations are explicitly derived by using the center manifold reduction and normal form theory. Finally, some numerical simulations are performed to illustrate and expand our theoretical results.     

Effect of prey‐taxis and diffusion on positive steady states for a predator‐prey system

In this paper, we consider a generalized predator‐prey system with prey‐taxis under the Neumann boundary condition. We investigate the local and global asymptotical stability of constant steady states (including trivial, semitrivial, and interior constant steady states). On the basis of a priori estimate and the fixed‐point index theory, several sufficient conditions for the nonexistence/existence of nonconstant positive solutions are given.     

Zero‐Hopf bifurcation in the Volterra‐Gause system of predator‐prey type

We prove that the Volterra‐Gause system of predator‐prey type exhibits 2 kinds of zero‐Hopf bifurcations for convenient values of their parameters. In the first, 1 periodic solution bifurcates from a zero‐Hopf equilibrium, and in the second, 4 periodic solutions bifurcate from another zero‐Hopf equilibrium. This study is done using the averaging theory of second order.     

Analysis of a nonautonomous eco‐epidemic diffusive model with disease in the prey

This paper deals with the behavior of positive solutions to a nonautonomous reaction‐diffusion system with homogeneous Neumann boundary conditions, which describes a two‐species predator‐prey system in which there is an infectious disease in prey. The sufficient condition on the permanence of the prey and the predator is established by combining the comparison principle with the results related to the corresponding ODE system. Some sufficient conditions for the spreading and vanishing of the disease are obtained. The global attractivity is also discussed by constructing a Lyapunov functional. Our results show that the disease is spreading if the transmission rate is suitably large, while if the transmission rate is small, the disease must be vanishing.     

Global bifurcation of solutions for a predator–prey model with prey‐taxis

We study pattern formations in a predator–prey model with prey‐taxis. It is proved that a branch of nonconstant solutions can bifurcate from the positive equilibrium only when the chemotactic is repulsive. Furthermore, we find the stable bifurcating solutions near the bifurcation point under suitable conditions. Copyright © 2014 John Wiley & Sons, Ltd.     

Persistence of two prey–one predator system with ratio‐dependent predator influence

In this study, we consider a mathematical model of two competing prey and one predator system where the prey species follow Lotka–Volterra‐type dynamics and the predator uptake functions are ratio dependent. We have derived the conditions for existence of different boundary equilibria and discussed their global behaviour. The sufficient condition for permanent co‐existence of all the species is derived. Finally, we have discussed the possibility of extinction of the species from the system. Copyright © 2000 John Wiley & Sons, Ltd.     

Complex dynamics of a discrete fractional‐order Leslie‐Gower predator‐prey model

A proposed discretized form of fractional‐order prey‐predator model is investigated. A sufficient condition for the solution of the discrete system to exist and to be unique is determined. Jury stability test is applied for studying stability of equilibrium points of the discretized system. Then, the effects of varying fractional order and other parameters of the systems on its dynamics are examined. The system undergoes Neimark‐Sacker and flip bifurcation under certain conditions. We observe that the model exhibits chaotic dynamics following stable states as the memory parameter α decreases and step size h increases. Theoretical results illustrate the rich dynamics and complexity of the model. Numerical simulation validates theoretical results and demonstrates the presence of rich dynamical behaviors include S‐asymptotically bounded periodic orbits, quasi‐periodicity, and chaos. The system exhibits a wide range of dynamical behaviors for fractional‐order α key parameter.     

A modified Leslie–Gower predator–prey model with ratio‐dependent functional response and alternative food for the predator

In this work, a modified Leslie–Gower predator–prey model is analyzed, considering an alternative food for the predator and a ratio‐dependent functional response to express the species interaction. The system is well defined in the entire first quadrant except at the origin ( 0 , 0 ) . Given the importance of the origin ( 0 , 0 ) as it represents the extinction of both populations, it is convenient to provide a continuous extension of the system to the origin. By changing variables and a time rescaling, we obtain a polynomial differential equations system, which is topologically equivalent to the original one, obtaining that the non‐hyperbolic equilibrium point ( 0 , 0 ) in the new system is a repellor for all parameter values. Therefore, our novel model presents a remarkable difference with other models using ratio‐dependent functional response. We establish conditions on the parameter values for the existence of up to two positive equilibrium points; when this happen, one of them is always a hyperbolic saddle point, and the other can be either an attractor or a repellor surrounded by at least one limit cycle. We also show the existence of a separatrix curve dividing the behavior of the trajectories in the phase plane. Moreover, we establish parameter sets for which a homoclinic curve exits, and we show the existence of saddle‐node bifurcation, Hopf bifurcation, Bogdanov–Takens bifurcation, and homoclinic bifurcation. An important feature in this model is that the prey population can go to extinction; meanwhile, population of predators can survive because of the consumption of alternative food in the absence of prey. In addition, the prey population can attain their carrying capacity level when predators go to extinction. We demonstrate that the solutions are non‐negatives and bounded (dissipativity and permanence of population in many other works). Furthermore, some simulations to reinforce our mathematical results are shown, and we further discuss their ecological meanings. Copyright © 2017 John Wiley & Sons, Ltd.     

PERSISTENCE AND STABILITY IN ARATIO‐DEPENDENT PREDATOR‐PREY SYSTEM WITH DELAY AND HARVESTING

ABSTRACT. . This paper aims to study the effect of time‐delay and combined harvesting on a Michaelis‐Menten type ratio‐dependent predator‐prey system. Dynamical behaviors such as persistence, stability, bifurcation, et cetera, are studied critically. Computer simulations are carried out to illustrate our analytical findings.