PPP项目中投资者协同行为发生机制及其演化规律研究

在公私合作项目(PPP)项目中,政府和私人投资者可能会采取协同行为来追求自身利益。这就需要对政府和私人投资者的协同行为进行研究,以了解提高项目绩效的基础机制。首先,基于演化博弈模型分析项目投资者策略选择的动态演化过程,据此政府和私人投资者通过交互选择来实现各自的最优策略。其次,通过演化博弈模型分析发现,政府和私人投资者协作管理具有多重复杂路径演化,其稳定策略很大程度上取决于组织的初始状态及相互激励关系。然后,探讨不同情境下投资者的最优策略和有效增强投资者协同行为的利益协调机制。     

Dynamics of a generalized Ricker–Beverton–Holt competition model subject to Allee effects

In this article, we propose and study a generalized Ricker–Beverton–Holt competition model subject to Allee effects to obtain insights on how the interplay of Allee effects and contest competition affects the persistence and the extinction of two competing species. By using the theory of monotone dynamics and the properties of critical curves for non-invertible maps, our analysis show that our model has relatively simple dynamics, i.e. almost every trajectory converges to a locally asymptotically stable equilibrium if the intensity of intra-specific competition intensity exceeds that of inter-specific competition. This equilibrium dynamics is also possible when the intensity of intra-specific competition intensity is less than that of inter-specific competition but under conditions that the maximum intrinsic growth rate of one species is not too large. The coexistence of two competing species occurs only if the system has four interior equilibria. We provide an approximation to the basins of the boundary attractors (i.e. the extinction of one or both species) where our results suggests that contest species are more prone to extinction than scramble ones are at low densities. In addition, in comparison to the dynamics of two species scramble competition models subject to Allee effects, our study suggests that (i) Both contest and scramble competition models can have only three boundary attractors without the coexistence equilibria, or four attractors among which only one is the persistent attractor, whereas scramble competition models may have the extinction of both species as its only attractor under certain conditions, i.e. the essential extinction of two species due to strong Allee effects; (ii) Scramble competition models like Ricker type models can have much more complicated dynamical structure of interior attractors than contest ones like Beverton–Holt type models have; and (iii) Scramble competition models like Ricker type competition models may be more likely to promote the coexistence of two species at low and high densities under certain conditions: At low densities, weak Allee effects decrease the fitness of resident species so that the other species is able to invade at its low densities; While at high densities, scramble competition can bring the current high population density to a lower population density but is above the Allee threshold in the next season, which may rescue a species that has essential extinction caused by strong Allee effects. Our results may have potential to be useful for conservation biology: For example, if one endangered species is facing essential extinction due to strong Allee effects, then we may rescue this species by bringing another competing species subject to scramble competition and Allee effects under certain conditions.     

Dynamics of an intraguild predation food web model with strong Allee effect in the basal prey

Since intraguild predation (IGP) is a ubiquitous and important community module in nature and Allee effect has strong impact on population dynamics, in this paper we propose a three-species IGP food web model consisted of the IG predator, IG prey and basal prey, in which the basal prey follows a logistic growth with strong Allee effect. We investigate the local and global dynamics of the model with emphasis on the impact of strong Allee effect. First, positivity and boundedness of solutions are studied. Then existence and stability of the boundary and interior equilibria are presented and the Hopf bifurcation curve at an interior equilibrium is given. The existence of a Hopf bifurcation curve indicates that if competition between the IG prey and IG predator for the basal resource lies below the curve then the interior equilibrium remains stable, while if it lies above the curve then the interior equilibrium loses its stability. In order to explore the impact of Allee effect, the parameter space is classified into sixteen different regions and, in each region, the number of interior equilibria is determined and the corresponding bifurcation diagrams on the Allee threshold are given. The extinction parameter regions of at least one species and the necessary coexistence parameter regions of all three species are provided. In addition, we explore possible dynamical patterns, i.e., the existence of multiple attractors. By theoretical analysis and numerical simulations, we show that the model can have one (i.e. extinction of all species), two (i.e. bi-stability) or three (i.e. tri-stability) attractors. It is also found by simulations that when there exists a unique stable interior equilibrium, the model may generate multiple attracting periodic orbits and the coexistence of all three species is enhanced as the competition between the IG prey and IG predator for the basal resource is close to the Hopf bifurcation curve from below. Our results indicate that the intraguild predation food web model exhibits rich and complex dynamic behaviors and strong Allee effect in the basal prey increases the extinction risk of not only the basal prey but also the IG prey or/and IG predator.     

三群体2×2×2非对称演化博弈稳定性分析

针对目前三方演化博弈的稳定性研究不足这一问题,利用复制动态方程构建了一般化的三维动力系统,首先讨论了单群体策略演化趋势,接着根据李雅普诺夫稳定性理论分析了系统的渐进稳定性,并结合单群体策略的演化趋势对系统稳定性作了深入研究。研究表明:严格纯策略纳什均衡是ESS,不严格纯策略纳什均衡是线性策略收敛(自定义概念),所有类型的混合策略纳什均衡均为鞍点,共同划分了ESS的吸引域,并证明了零特征值非ESS定理,以及ESS不共边定理,在此基础上给出了N维双策略系统中ESS的最多个数。最后,设计了六组经典算例,首先结合研究结果分析了算例,接着对算例进行系统仿真,仿真结果与理论分析一致,为演化博弈的进一步研究提供借鉴与启发。     

Adaptive Evolution Analysis of a Predator-Prey Model With Group Defense北大核心CSCD

Based on the theoretical framework of adaptive dynamics, the evolution of the predator-prey model with functional response of group defense effect on the predator handling time, was investigated. Firstly, in view of the interaction of predator populations with interspecific competition, the evolutionary conditions for a single predator population to split into 2 populations with different strategies through evolutionary branching were given. Secondly, when the ecological equilibrium of the model is unstable and the system has a limit cycle, the population will have strong coexistence under large mutation, but this coexistence will be evolutionarily unstable. Finally, the conclusions for the model with Holling-Ⅱ type functional response were compared. The results indicate that, with a sufficiently large prey carrying capacity, group defense effects can evolutionarily lead to the extinction of predators. © 2023 Editorial Office of Applied Mathematics and Mechanics. All rights reserved.     

Multiple stability and uniqueness of the limit cycle in a Gause-type predator–prey model considering the Allee effect on prey

In this work, a bidimensional differential equation system obtained by modifying the well-known predator–prey Rosenzweig–MacArthur model is analyzed by considering prey growth influenced by the Allee effect.One of the main consequences of this modification is a separatrix curve that appears in the phase plane, dividing the behavior of the trajectories. The results show that the equilibrium in the origin is an attractor for any set of parameters. The unique positive equilibrium, when it exists, can be either an attractor or a repeller surrounded by a limit cycle, whose uniqueness is established by calculating the Lyapunov quantities. Therefore, both populations could either reach deterministic extinction or long-term deterministic coexistence.The existence of a heteroclinic curve is also proved. When this curve is broken by changing parameter values, then the origin turns out to be an attractor for all orbits in the phase plane. This implies that there are plausible conditions where both populations can go to extinction. We conclude that strong and weak Allee effects on prey population exert similar influences on the predator–prey model, thereby increasing the risk of ecological extinction.     

Weak Allee effects and species coexistence

In this article, we study the population dynamics of a two-species discrete-time competition model where each species suffers from either predator saturation induced Allee effects and/or mate limitation induced Allee effects. We focus on the following two possible outcomes of the competition: 1. one species goes to extinction; 2. the system is permanent. Our results indicate that, even if one species’ intra-specific competition is less than its inter-specific competition, weak Allee effects induced by predation saturation can promote coexistence of the two competing species. This is supported by the outcome of two-species competition models without Allee effects. Also, we discuss our results and future work on multiple attractors in competition models with Allee effects.     

Evolutionary analysis of adaptive dynamics model under variation of noise environment

This paper proposes a stochastic model for the evolutionary adaptive dynamics of species subject to trait-dependent intrinsic growth rates and the influence of environmental noise. The aim of this paper is twofold: (a) mathematically we make an attempt to investigate the evolutionary adaptive dynamics for models with noises; (b) biologically we investigate how the noises in environment affect the evolutionary stability. We first investigate the extinction and permanence of the population in the presence of environmental noises. Combining evolutionary adaptive dynamics with stochastic dynamics, we then establish a fitness function with stochastic disturbance and obtain the evolutionary conditions for continuously stable strategy and evolutionary branching. Our study finds that under intense competition among species, increasing stochastic disturbance can lead to rapidly stable evolution towards smaller trait values, but there is an opposite effect under weak competition among species. This yields an interesting evolutionary threshold, beyond which any increasing stochastic disturbance can go against evolutionary branching and promote evolutionary stability. We then carry out the evolutionary analysis and numerical simulations to illustrate our theoretical results. Finally, for demonstrating the emergence of high-level polymorphism we perform long-term simulation of evolutionary dynamics.     

基于强互惠理论的企业合作创新行为演化研究

基于强互惠理论构建了企业合作创新行为演化模型,将企业分成投机、纯合作与强互惠三种类型,对基本的演化博弈模型进行了分析,利用复制动态方程求出基本模型的两个演化稳定均衡解,并通过仿真方法对放松约束后的演化模型做进一步分析,发现了新的系统均衡状态并给出解释。研究结果表明:强互惠行为可以通过演化生成,并抑制企业合作创新中的投机行为,与其他两种类型企业共同发展,并形成企业合作创新系统的演化稳定均衡。研究结果为企业合作创新组织的管理提供了决策依据,同时有助于推动企业合作创新理论的深入发展。     

A global bifurcation theorem for Darwinian matrix models

Motivated by models from evolutionary population dynamics, we study a general class of nonlinear difference equations called matrix models. Under the assumption that the projection matrix is non-negative and irreducible, we prove a theorem that establishes the global existence of a continuum with positive equilibria that bifurcates from an extinction equilibrium at a value of a model parameter at which the extinction equilibrium destabilizes. We give criteria for the global shape of the continuum, including local direction of bifurcation and its relationship to the local stability of the bifurcating positive equilibria. We discuss a relationship between backward bifurcations and Allee effects. Illustrative examples are given.     

Colony and evolutionary dynamics of a two‐stage model with brood cannibalism and division of labor in social insects

Division of labor (DOL) is a major factor for the great success of social insects because it increases the efficiency of a social group where different individuals perform different tasks repeatedly and presumably with increased performance. Cannibalism plays an important role in regulating colony growth and development by regulating the number of individuals in a colony and increasing survival by providing access to essential nutrients and minimizing competition among colony mates. To understand the synergy effects of DOL and cannibalistic behavior on colony dynamic outcomes, we propose and study a compartmental two‐stage model using ecological and evolutionary game theory settings. Our analytical results of the ecological and evolutionary models suggest that: (1) A noncannibalistic colony can survive if the efficiency of energy investment reflecting the DOL is greater than the relative death rate of the older population. (2) A cannibalistic colony can die out if both the efficiency of energy investment and the relative cannibalism rate (where each is also reflecting the DOL) are too large; or if the relative cannibalism rate alone is too small. (3) From our numerical analysis, cannibalism can increase or reduce the colony's total population size, which greatly depends on the benefit of egg cannibalism increasing or decreasing of adult's lifespan. (4) A cannibalistic and noncannibalistic colony can experience bistability due to cooperative behavior. (5) In the evolutionary settings, DOL can prevent colony death and natural selection can preserve strong Allee effects by selecting the traits with the largest investment on brood care and the lowest cannibalism rate. (6) Evolutionary dynamics may increase the fitness of the colony, i.e., the successful production of workforce which results in the increase of total worker population size, colony survival, and reproduction. Our results suggest both cannibalism and DOLs are adaptive strategies that increase the size of the worker population, and therefore, persistence of the colony.     

具有HollingⅡ型功能反应和Allee效应的捕食系统模型

利用计算机模拟方法研究一类离散种群相互作用模型的动态复杂性.通过理论推导建立食饵具有Allee效应和HollingⅡ型功能反应的自治捕食系统模型,用Matlab软件模拟离散种群的生长状态,探索研究参数的变化对种群大小的影响,阐释Allee效应及HollingⅡ型功能反应在种群间相互作用模型中的重要性.研究结果表明:1)当处理时间处于有效区间内时,处理时间越大种群的稳定共存参数域越大;2)Allee效应的引入使种群的动态行为更为复杂,从而增加了捕食者种群的灭绝风险;3)系统受强Allee效应的影响,种群会出现提前分叉现象,如果继续增加Allee效应就会导致种群灭绝;4)强Allee效应更容易使种群趋向灭绝.所得结论在丰富生态学理论的同时,提出了保护生态学的重要依据.     

The impact of Allee effect on a predator–prey system with Holling type II functional response

In this paper, the Allee effect is incorporated into a predator–prey model with Holling type II functional response. Compared with the predator–prey model without Allee effect, we find that the Allee effect of prey species increases the extinction risk of both predators and prey. When the handling time of predators is relatively short and the Allee effect of prey species becomes strong, both predators and prey may become extinct. Moreover, it is shown that the model with Allee effect undergoes the Hopf bifurcation and heteroclinic bifurcation. The Allee effect of prey species can lead to unstable periodical oscillation. It is also found that the positive equilibrium of the model could change from stable to unstable, and then to stable when the strength of Allee effect or the handling time of predators increases continuously from zero, that is, the model admits stability switches as a parameter changes. When the Allee effect of prey species becomes strong, longer handling time of predators may stabilize the coexistent steady state.     

Mathematical models of restoration and control of a single species with Allee effect

According to the initial density of a single species with Allee effect and corresponding management strategy, three kinds of mathematical models are presented to describe the evolutionary process of the species by impulsive differential equations. When the initial density of the species is larger than economic injury level (EIL) (or economical threshold, ET), impulsive harvest control is considered in a finite time to decrease the population of the species. The feasibility of the impulsive harvest control in a finite time is given by the existence of solution of the model with initial and boundary value problem. When the initial density of the species is less than EIL (or ET), the model with state feedback control is formulated according to the state of the species. The existence and stability of periodic solution of the model with state feedback control are discussed. When the initial density of the species is less than the Allee threshold and the species tends to extinction, the model with impulsive release at fixed moments is presented to study the restoration of the species. The conditions for the feasibility of periodic restoration of the species are given. Finally, some discussions are given.     

Complex dynamics of a discrete predator–prey model with the prey subject to the Allee effect

In this paper, complex dynamics of the discrete predator–prey model with the prey subject to the Allee effect are investigated in detail. Firstly, when the prey intrinsic growth rate is not large, the basins of attraction of the equilibrium points of the single population model are given. Secondly, rigorous results on the existence and stability of the equilibrium points of the model are derived, especially, by analyzing the higher order terms, we obtain that the non-hyperbolic extinction equilibrium point is locally asymptotically stable. The existences and bifurcation directions for the flip bifurcation, the Neimark–Sacker bifurcation and codimension-two bifurcations with 1:2 resonance are derived by using the center manifold theorem and the bifurcation theory. We derive that the model only exhibits a supercritical flip bifurcation and it is possible for the model to exhibit a supercritical or subcritical Neimark–Sacker bifurcation at the larger positive equilibrium point. Chaos in the sense of Marotto is proved by analytical methods. Finally, numerical simulations including bifurcation diagrams, phase portraits, sensitivity dependence on the initial values, Lyapunov exponents display new and rich dynamical behaviour. The analytic results and numerical simulations demonstrate that the Allee effect plays a very important role for dynamical behaviour.     

基于演化博弈的发电侧电力市场长期均衡模型

研究包含生产同质电力产品的两组 (种群 )企业——低成本发电企业和高成本发电企业的发电侧电力市场的长期均衡问题 .应用演化博弈论的有限种群演化稳定战略概念 ,证明了有限种群的演化稳定战略产量分别等于两组 (种群 )企业的竞争产量 .通过建立基于企业战略模仿和试验的随机演化模型 ,分析了发电侧电力市场长期均衡的演化过程 .     

On chaos, transient chaos and ghosts in single population models with Allee effects

Density-dependent effects, both positive or negative, can have an important impact on the population dynamics of species by modifying their population per-capita growth rates. An important type of such density-dependent factors is given by the so-called Allee effects, widely studied in theoretical and field population biology. In this study, we analyze two discrete single population models with overcompensating density-dependence and Allee effects due to predator saturation and mating limitation using symbolic dynamics theory. We focus on the scenarios of persistence and bistability, in which the species dynamics can be chaotic. For the chaotic regimes, we compute the topological entropy as well as the Lyapunov exponent under ecological key parameters and different initial conditions. We also provide co-dimension two bifurcation diagrams for both systems computing the periods of the orbits, also characterizing the period-ordering routes toward the boundary crisis responsible for species extinction via transient chaos. Our results show that the topological entropy increases as we approach to the parametric regions involving transient chaos, being maximum when the full shift R(L)^∞ occurs, and the system enters into the essential extinction regime. Finally, we characterize analytically, using a complex variable approach, and numerically the inverse square-root scaling law arising in the vicinity of a saddle-node bifurcation responsible for the extinction scenario in the two studied models. The results are discussed in the context of species fragility under differential Allee effects.     

新零售背景下生鲜超市的演化博弈分析

本文聚焦当前新零售背景下生鲜超市的竞争态势,借助于演化博弈这一研究工具,基于经济学原理构建了生鲜电商切入线下生鲜超市以后,与传统生鲜超市为竞争生存空间而形成的演化博弈模型,通过模型分析探讨这一演化过程中的演化逻辑,发现其中的演化规律及关键影响因素。研究认为,虽然打着“新零售”旗号的电商平台在切入线下生鲜超市时,通过首选精品化策略可以在初始阶段实现与传统生鲜超市的差异化竞争,但是,双方能否就此达成稳定均衡将会受到多种因素的制约,并可能演化出完全不同的均衡状态;甚至在特定的市场条件下,精品化都未必是生鲜超市的理性选择。     

Transitions in a logistic population model incorporating an Allee effect

In some species, the population may decline to zero; that is, the species becomes extinct if the population falls below a given threshold. This phenomenon is well known as an Allee effect. In most Allee models, the model parameters are constants, and the population tends either to a nonzero limiting state (survival) or to zero (extinction). However, when environmental changes occur, these parameters may be slowly varying functions of time. Then, application of multitiming techniques allows us to construct approximations to the evolving population in cases where the population survives to a slowly varying surviving state and those where the population declines to zero. Here, we investigate the solution of a logistic population model exhibiting an Allee effect, when the carrying capacity and the limiting density interchange roles, via a transition point. We combine multiscaling analysis with local asymptotic analysis at the transition point to obtain an overall expression for the evolution of the population. We show that this shows excellent agreement with the results of numerical computations. Copyright © 2015 John Wiley & Sons, Ltd.